Figure 1: Paired black-tailed godwits arrive at their breeding sites synchronously but winter separately.

a, The arrival dates (days from 31 March) of paired males and females were highly correlated (r = 0.97, P<0.001); the regression line indicates identical arrival times for males and females. For each pair, data are for one year only. b, Wintering locations of pairs of godwits. Each line links a pair and the circle indicates the female's location. Locations of 12 of the 14 pairs are known in the same winter. Godwits show a very high degree of between-year philopatry in winter.