Figure 1: Coupling between catalysis and rotation in F1.
From: ATP hydrolysis assists phosphate release and promotes reaction ordering in F1-ATPase

Dwells in the γ rotation and the corresponding nucleotide states at each β-subunit (coloured circles) are shown. Nucleotides are coloured according to the β-subunits for visual clarity. The grey arrows represent the rotary angles of the γ-subunit. ATP binding, hydrolysis, ADP release and Pi release of the red β-subunit occur at 0°, 200°, 240° and 320°, respectively.