Fig. 2: DCAP blocks the autophagic flux without impairing mitochondrial potential.

a–e Immunoblot analysis of LC3, SQSTM1, AMPK phosphorylated in T172 (pAMPK) and ULK1 phosphorylated in S555 (pULK1) in U2OS cells treated 24 h with three increasing doses of DCAP (1, 5 and 10 μM) or CCCP (5, 10 and 20 μM). GAPDH was used as a loading control. Densitometry analysis of LC3-II (b), SQSTM1 (c), pAMPK (d) and pULK1 (e) is reported as relative protein levels normalized by GAPDH. Vehicle (DMSO) sample value was set to 1 (dotted lines in the graphs). Shown as mean ± s.e.m., n = 3. *P < 0.05; **P < 0.01; ***P < 0.001 DCAP versus CCCP (two-way ANOVA with Bonferroni post test). f Loss of mitochondrial membrane potential (ΔΨm) was quantified by measuring the amount of monomeric JC-10 dye after 24 h of treatment with vehicle (DMSO), 10 µM DCAP or 20 µM CCCP. These doses were adopted as they produced a comparable LC3-II induction (b). Shown as mean ± s.e.m., n = 6. ***P < 0.001, CCCP versus vehicle (one-way ANOVA). g Immunoblot analysis of cytosolic and organelles-enriched (vacuolar) preparations from cells treated 24 h with vehicle (DMSO), 10 µM DCAP or 25 µM chloroquine (CQ). Autophagic LC3-II and lysosomal LAMP1 proteins were used to confirm the enrichment in autophagosomes and autophagolysosomes in vacuolar fractions. Cytoplasmic TUBULIN protein was adopted to evaluate potential cytosolic contaminations in the organelles-enriched fraction. Immunoblot with antibodies against the autophagic receptors, SQSTM1 and NBR1, showed accumulation of these proteins in vacuolar fractions of DCAP- and CQ-treated cells. h Autophagosomal degradation of a SQSTM1 protein fused with a Red Fluorescent Protein (RFP) was observed by live fluorescent microscopy in SQSTM1-RFP expressing cells treated with DMSO, 10 µM DCAP or 25 µM CQ. Representative images before addition of compound (time 0) and at 6, 12, and 24 h post-treatment are shown. DCAP and CQ treatment produced accumulation of SQSTM1-fluorescent dots, overtime. Scale bar in the 24 h frame = 5 µm