Fig. 1

Effect of background climate on LU and GDDreq. Relative importance (%) of each variable in estimating (a) observed LU date for all sites and species (n = 27790), (b) GDDreq for all sites and species, (c) LU date for drought-prone sites (n = 5747) and (d) GDDreq for drought-prone sites. See the last row of Fig. 2a–d (ALL) for the direction of each effect. NCD, number of chilling days estimated as the number of days between 1 November in the previous year and the LU date with temperatures between 0 and 5 °C; TG, mean growing-season temperature; TP, mean pre-season temperature; SWG, mean growing-season shortwave [visible and near infrared] radiation; LWG, mean growing-season longwave [infrared] radiation; SWP, mean pre-season shortwave [visible and near infrared] radiation; LWP, mean pre-season longwave [infrared] radiation; PG growing-season total precipitation; PP, pre-season total precipitation; DL, day length at LU date; SMG, growing season soil-moisture content; SMP, pre-season soil-moisture content αE, ratio of actual to potential evapotranspiration. drought-prone sites were defined as sites with long-term αE <0.9. Variables were selected based on penalized elastic net regression and corrected for spatial autocorrelation; the coefficient of determination (R²) of the selected models is given at the top of each panel. Error bars represent the 95% confidence interval. The sum of the relative importance equals 100% of R². The scatter plot in each panel represents modeled versus observed variables (LU or GDDreq) after correcting coefficients for collinearity and spatial autocorrelation. The red line represents the 1:1 line. Source data are provided as a Source Data file.