Fig. 1: Anatomical overview of Youti yuanshi.

YKLP 12387. a, External scanning electron microscopy, right side. Damage to posterior epidermis exposes lining of perivisceral cavity, demonstrating blind gut. b, External scanning electron microscopy, left side. c,g–j, Median virtual dissection from X-ray computed tomography (XCT) data (c), showing location of transverse slices intersecting digestive glands (g,i) and transverse membrane (h,j). d, Semi-manual segmentation of internal chambers from XCT data, viewed from the left side. Dorsolateral aspects of the peripheral cavity are omitted for clarity. e,f, Virtual dissection parallel to coronal plane, looking ventrally (e) and dorsally (f), showing digestive glands, pericardial sinus, transverse membranes within perivisceral cavity, and oblique membranes within peripheral cavity. g–j, XCT sections at positions indicated in c at position of digestive glands (g,i) and at position of ventrolateral lacunae and transverse membrane (h,j). g,h, Sections close to the anterior trunk, reflecting segments at late developmental stage. i,j, Sections close to the posterior trunk, showing superior preservation of internal tissue. k, Segmentation of internal chambers from XCT data, viewed from the dorsal perspective at anterior, middle and posterior trunk. Aspects of peripheral cavity are omitted for clarity. a, appendage; cb, central body of brain; db, dorsolateral body of brain; dia, diagenetic grain; dg, digestive gland; dm, dorsal membrane; dp, dorsal projection; dv, dorsal vessel; fb, frontal body of brain; irr, irregular chamber; lig, ligament; om, oblique membrane; pc, pericardial sinus; pph, peripheral cavity; pn, perineural sinus; pv, perivisceral cavity; tm, transverse membrane; vl, ventrolateral sinus; vv, ventral vessel. Scale bars, 200 μm.