Table 2 The summary of morphological characters of the male imagines to distinguish extant and extinct representatives of the genus Calliarcys. Distinct differential characters are marked in bold.

From: Cenozoic origins of the genus Calliarcys (Insecta, Ephemeroptera) revealed by Micro-CT, with DNA barcode gap analysis of Leptophlebiinae and Habrophlebiinae

Characters (male imago)

Calliarcys antiquus sp. nov.

Calliarcys humilis

Calliarcys van

Head

   

Compound eyes colour: upper portion

Creamy to orange pink

Brick-red

Compound eyes colour: lower portion

Black to blackish grey

Black to blackish grey

Compound eyes: shape

Distinctly large, contiguous

Middle-sized, widely separated

Large, separated

Wings: cross venation

Well-developed

Ordinarily developed

Ordinarily developed

Forewing

   

Width/length ratio

0.38

0.35–0.38

0.36–0.38

Pterostigmatic venation

6–7 forked and 2–3 simple veins

8–12 simple veins

6–8 simple veins

Cubital venation: longest vein

iCu2

iCu3

iCu3

Hind wing

   

Ratio to forewing length

3.74–3.81

3.30–3.60

3.8–4.0

Width/length ratio

0.54

0.50–0.54

0.52–0.54

Costal process: location

Middle of wing length

Before middle of length

Middle of wing length

Costal process: shape

Widely rounded apically

Step-like, acute apically

Smoothly rounded apically

Number of cross veins between MA and MP

8

1–2

1–2

Number of cross veins between MP and CuA

2

0

0

Cubital triad

Complete

Not complete

Not complete

Abdomen

   

Colour of terga

Uniformly brown to dark brown

Yellowish-brown strip anteriorly

Genitalia

   

Median projections of styliger: shape

Nearly finger-like

Finger-like

Nearly triangular

Forceps segment I: inner hump

Present

Absent

Absent

I. Penis lobes: general shape

(?) separated basally

Fussed basally

Fussed basally

II. Penis lobes: general shape

Relatively robust, shortened

Relatively elongated

Relatively robust, shortened

III. Penis lobes: shape of tip

Slightly bent inward

Distinctly bent inward

Slightly bent inward