Abstract
The ability of early hominins to adapt to diverse landscapes and ecological niches was a key factor in their range expansion. However, the behavioral and cognitive prerequisites underlying this adaptability remain poorly understood. Assuming that resource procurement played a pivotal role in hominin dispersal, we investigate the mobility-subsistence system of early Oldowan groups who occupied the Lower Omo landscape of southern Ethiopia some 2.3 million years ago (Ma). With its extensive archaeological record and scattered lithic resources, this context provides a landscape-scale record of early hominin behavior. Multiple lines of evidence derived from an integrated environmental, spatial and technological analysis indicate a regionally-structured mobility strategy, involving the provisioning of quartz pebbles and the transport of partially transformed products from distant sources to the biotic-rich but stone-poor margins of the paleo-Omo River, where highly mobile foraging activities were concentrated along the meandering channel. This regional strategy of early hominin displacement reflects a “Shungura exception” in the Early Pleistocene record. It coincides with the emergence of stone tool-mediated activities in the Lower Omo Valley, marking a tipping point in the capacity of early hominins to develop environment-specific adaptive strategies‒a crucial asset for their expansion into new ecological niches.
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All data generated or analyzed during this study are included in this published article and its supplementary information file, or publicly accessible through Nakala data repository (https://nakala.fr/10.34847/nkl.25d73am0).
References
Kuhn, S. L., Raichlen, D. A. & Clark, A. E. What moves us? How mobility and movement are at the center of human evolution. Evol. Anthropol. Issues News Rev. 25, 86–97 (2016).
Nathan, R. et al. A movement ecology paradigm for unifying organismal movement research. Proc. Natl. Acad. Sci. 105, 19052–19059 (2008).
Potts, R. Environmental and behavioral evidence pertaining to the evolution of early homo. Curr. Anthropol. 53, S299–S318 (2012).
Gabunia, L. & Vekua, A. A plio-pleistocene hominid from Dmanisi, East Georgia, Caucasus. Nature 373, 509–512 (1995).
Mgeladze, A. et al. Hominin occupations at the Dmanisi site, Georgia, Southern caucasus: Raw materials and technical behaviours of europe’s first hominins. J. Hum. Evol. 60, 571–596 (2011).
Harmand, S. et al. 3.3-million-year-old stone tools from Lomekwi 3, West Turkana, Kenya. Nature 521, 310–315 (2015).
Semaw, S. et al. 2.5-million-year-old stone tools from gona. Ethiopia Nature. 385, 333–336 (1997).
Plummer, T. W. et al. Expanded geographic distribution and dietary strategies of the earliest Oldowan hominins and Paranthropus. Science 379, 561–566 (2023).
Braun, D. R. et al. Earliest known Oldowan artifacts at > 2.58 Ma from Ledi-Geraru, Ethiopia, highlight early technological diversity. Proc. Natl. Acad. Sci. 116, 11712–11717 (2019).
Roche, H. Hadar et les industries préacheuléennes d’afrique orientale. Bull. Société Préhistorique Fr. 79, 165–166 (1982).
Roche, H. et al. Early hominid stone tool production and technical skill 2.34 Myr ago in West Turkana, Kenya. Nature 399, 57–60 (1999).
Kimbel, W. H. et al. Late pliocene homo and Oldowan tools from the Hadar formation (Kada Hadar Member), Ethiopia. J. Hum. Evol. 31, 549–567 (1996).
Chavaillon, J. Découverte d’un Niveau Oldowayen Dans La Basse vallée de l’omo. Bull. Société Préhistorique Fr. 67, 7–11 (1970).
Merrick, H. V., De Heinzelin, J., Haesaerts, P. & Howell, F. C. Archaeological occurences of early pleistocene age from the Shungura Formation, lower Omo Valley, Ethiopia. Nature 242, 572–575 (1973).
de Heinzelin, J. et al. Environment and behavior of 2.5-million-year-old bouri hominids. Science 284, 625–629 (1999).
Rogers, M. J., Harris, J. W. & Feibel, C. S. Changing patterns of land use by Plio-Pleistocene hominids in the Lake Turkana basin. J. Hum. Evol. 27, 139–158 (1994).
Plummer, T. Flaked stones and old bones: biological and cultural evolution at the dawn of technology. Am. J. Phys. Anthropol. 125, 118–164 (2004).
Stout, D., Quade, J., Semaw, S., Rogers, M. J. & Levin, N. E. Raw material selectivity of the earliest stone toolmakers at Gona, Afar, Ethiopia. J. Hum. Evol. 48, 365–380 (2005).
Goldman-Neuman, T. & Hovers, E. Raw material selectivity in late pliocene Oldowan sites in the Makaamitalu Basin, Hadar, Ethiopia. J. Hum. Evol. 62, 353–366 (2012).
Harmand, S. Variability in Raw material selectivity at the late pliocene sites of Lokalalei, West Turkana, Kenya. in Interdisciplinary Approaches To the Oldowan (eds (eds Hovers, E. & Braun, D. R.) 85–97 (Springer, Dordrecht, (2009).
Delagnes, A. et al. Archaeological investigations in the lower Omo Valley (Shungura Formation, Ethiopia): New data and perspectives. J. Hum. Evol. 61, 215–222 (2011).
Delagnes, A. & Roche, H. Late pliocene hominid knapping skills: The case of Lokalalei 2 C, West Turkana, Kenya. J. Hum. Evol. 48, 435–472 (2005).
Stout, D., Semaw, S., Rogers, M. J. & Cauche, D. Technological variation in the earliest Oldowan from Gona, Afar, Ethiopia. J. Hum. Evol. 58, 474–491 (2010).
Potts, R. Why the oldowan? Plio-pleistocene tool-making and the transport of resources. J. Anthropol. Res. 47, 153–176 (1991).
Braun, D. R. et al. Oldowan behavior and raw material transport: Perspectives from the Kanjera formation. J. Archaeol. Sci. 35, 2329–2345 (2008).
Favreau, J. Sourcing Oldowan and Acheulean stone tools in Eastern Africa: Aims, methods, challenges, and state of knowledge. Quat Sci. Adv. 9, 100068 (2023).
Blumenschine, R. J. et al. Late pliocene homo and hominid land use from Western Olduvai Gorge, Tanzania. Science 299, 1217–1221 (2003).
Stiles, D. Early hominid behaviour and culture tradition: Raw material studies in Bed II, Olduvai gorge. Afr. Archaeol. Rev. 9, 1–19 (1991).
Delagnes, A. et al. Long-term behavioral adaptation of Oldowan toolmakers to resource-constrained environments at 2.3 Ma in the Lower Omo Valley (Ethiopia). Sci. Rep. 13, 14350 (2023).
McDougall, I. & Brown, F. H. Geochronology of the pre-KBS tuff sequence, Omo Group, Turkana basin. J. Geol. Soc. 165, 549–562 (2008).
McDougall, I. et al. New single crystal 40Ar/39Ar ages improve time scale for deposition of the Omo Group, Omo–Turkana Basin, East Africa. J. Geol. Soc. 169, 213–226 (2012).
de Heinzelin, J. The Omo Group. Archives of the International Omo Research Expedition Vol. 85 (Musée Royal de l’Afrique Centrale, 1983).
de Heinzelin, J. & Haesaerts, P. Plio-Pleistocene formations of the lower Omo basin with particular reference to the Shungura formation. in Earliest Man and Environments in the Lake Rudolf Basin (eds Coppens, Y., Howell, F. C. & Isaac, G.,L. & Leakey, R. E. F.) 24–49 (Chicago, 1976).
Braun, D. R., Harris, J. W. & Maina, D. Oldowan raw material procurement and use: Evidence from the Koobi fora formation. Archaeometry 51, 26–42 (2009).
Tiercelin, J. J. et al. New considerations on the stratigraphy and environmental context of the oldest (2.34 Ma) Lokalalei archaeological site complex of the Nachukui Formation, West Turkana, Northern Kenya rift. J. Afr. Earth Sci. 58, 157–184 (2010).
Howell, F. C., Haesaerts, P. & de Heinzelin, J. Depositional environments, archeological occurrences and hominids from member F and E of the Shungura formation (Omo Basin, Ethiopia). J. Hum. Evol. 16, 665–700 (1987).
Barton, C. M. & Riel-Salvatore, J. The fomation of lithic assemblages. J. Archaeol. Sci. 46, 334–352 (2014).
Chavaillon, J. Evidence for the technical practices of early pleistocene hominids, Shungura Formation, lower Omo Valley, Ethiopia. in Earliest Man and Environments in the Lake Rudolf Basin (eds (eds Coppens, Y., Howell, F. C., Isaac, G. L. I. & Leakey, R.) 565–573 (University of Chicago Press, Chicago and London, (1976).
Chavaillon, J. & Boisaubert, J. L. Prospection archéologique Dans Le Gemu-Gofa et La Basse vallée de l’omo. Abbay Cahier n°8 3–10 (1977).
Rothman, K. J. No adjustements are needed for multiple comparisons. Epidemiology 1, 43–46 (1990).
Escoufier, Y. Le traitement des variables vectorielles. Biometrics 29, 751 (1973).
Delagnes, A., Brenet, M., Gravina, B. & Santos, F. Exploring the relative influence of raw materials, percussion techniques, and hominin skill levels on the diversity of the early Oldowan assemblages: Insights from the Shungura Formation, lower Omo Valley, Ethiopia. PLOS ONE. 18, e0283250 (2023).
Hovers, E. & Braun, D. R. Interdisciplinary Approaches To the Oldowan (Springer, 2009).
de la Torre, I. Omo revisited. Evaluating the technological skills of pliocene hominids. Curr. Anthropol. 45, 439–465 (2004).
Toth, N. The Oldowan reassessed: A close look at early stone artifacts. J. Archaeol. Sci. 12, 101–120 (1985).
Turq, A., Roebroeks, W., Bourguignon, L. & Faivre, J. P. The fragmented character of middle palaeolithic stone tool technology. J Hum. Evol. 641–655 (2013).
Geneste, J. M. L’approvisionnement En matières premières Dans les systèmes de production lithique: La dimension spatiale de La technologie. in Tecnologia Y Cadenas Operativas Liticas (eds (eds Mora, R., Terradas, X., Parpal, A. & Plana, C.) 1–36 (Departament d’Historia de les Societats Pre-capitalistes i d’Antropologia Social, Bellaterra, (1991).
Toth, N. Behavioral inferences from early stone artifact assemblages: An experimental model. J. Hum. Evol. 16, 763–787 (1987).
Tavoso, A. Réflexions Sur l’économie des matières premières Au Moustérien. Bull. Société Préhistorique Fr. 81, 79–82 (1984).
Tavoso, A. Les industries de La moyenne Terrasse du Tarn à Tecou (Tarn). Bull. Mus. Anthropol. Préhistorique Monaco. 18, 113–144 (1972).
Merrick, H. V. Recent archaeological research in the Plio-Pleistocene deposits of the lower Omo, Southwestern Ethiopia. Hum Orig Louis Leakey East. Afr. Evid. 461–481 (1976).
Ludwig, B. V. A Technological Reassessment of East African Plio-Pleistocene Lithic Artifact assemblages. vol. PhD (The State University of New Jersey, 1999).
Barsky, D. et al. The early Oldowan stone-tool assemblage from Fejej FJ-1A, Ethiopia. J. Afr. Archaeol. 9, 207–224 (2011).
Abrunhosa, A. et al. Understanding Neanderthal technological adaptation at Navalmaíllo Rock Shelter (Spain) by measuring lithic raw materials performance variability. Archaeol. Anthropol. Sci. 11, 5949–5962 (2019).
Key, A., Proffitt, T. & De La Torre, I. Raw material optimization and stone tool engineering in the early stone age of Olduvai Gorge (Tanzania). J. R Soc. Interface. 17, 20190377 (2020).
Pargeter, J. & de la Peña, P. Milky quartz bipolar reduction and lithic miniaturization: Experimental results and archaeological implications. J. Field Archaeol. 42, 551–565 (2017).
Mourre, V. Les industries En quartz Au Paléolithique. Terminologie, méthodologie et technologie. Paléo Rev. Archéologie Préhistorique. 8, 205–223 (1996).
Tallavaara, M., Manninen, M. A., Hertell, E. & Rankama, T. How flakes shatter: A critical evaluation of quartz fracture analysis. J. Archaeol. Sci. 37, 2442–2448 (2010).
Galland, A. Préservation Et Fonction Des Outillages De l’Oldowayen Ancien: Application Au Registre Lithique Du Membre F De La Formation De Shungura (Basse Vallée De l’Omo, Éthiopie) (Université de Bordeaux, 2022).
Blumenschine, R. J., Masao, F. T., Tactikos, J. C. & Ebert, J. I. Effects of distance from stone source on landscape-scale variation in Oldowan artifact assemblages in the Paleo-Olduvai Basin, Tanzania. J. Archaeol. Sci. 35, 76–86 (2008).
Nash, D. J. et al. Provenancing of silcrete Raw materials indicates long-distance transport to Tsodilo Hills, Botswana, during the middle stone age. J. Hum. Evol. 64, 280–288 (2013).
Will, M. The role of different Raw materials in lithic technology and settlement patterns during the middle stone age of Southern Africa. Afr. Archaeol. Rev. 38, 477–500 (2021).
Kuhn, S. L. Mousterian Lithic Technonlogy: An Ecological Perspective (Princeton University Press, 1995).
Barr, W. A. Paleoenvironments of the Shungura formation (Plio-Pleistocene: Ethiopia) based on ecomorphology of the Bovid astragalus. J. Hum. Evol. 88, 97–107 (2015).
Sept, J. M. Beyond bones: Archaeological sites, early hominid subsistence, and the costs and benefits of exploiting wild plant foods in East African riverine landscapes. J. Hum. Evol. 27, 295–320 (1994).
Reynolds, S. C., Bailey, G. N. & King, G. C. Landscapes and their relation to hominin habitats: Case studies from australopithecus sites in Eastern and Southern Africa. J. Hum. Evol. 60, 281–298 (2011).
Bobe, R. & Eck, G. G. Responses of African Bovids to pliocene Climatic change. Paleobiology 27, 1–48 (2001).
Alemseged, Z. An integrated approach to taphonomy and faunal change in the Shungura formation (Ethiopia) and its implication for hominid evolution. J. Hum. Evol. 44, 451–478 (2003).
Negash, E. W. et al. Dietary trends in herbivores from the Shungura Formation, Southwestern Ethiopia. Proc. Natl. Acad. Sci. 117, 21921–21927 (2020).
Gardin, A. et al. Stable oxygen isotopes of crocodilian tooth enamel allow tracking Plio-Pleistocene evolution of freshwater environments and climate in the Shungura formation (Turkana Depression, Ethiopia). Biogeosciences 21, 437–454 (2024).
Linares-Matás, G. J. & Clark, J. Seasonality and Oldowan behavioral variability in East Africa. J. Hum. Evol. 164, 103070 (2022).
Clark, J. & Linares-Matás, G. J. Seasonality and lithic investment in the Oldowan. J. Paleolit Archaeol. 6, 38 (2023).
Alemseged, Z., Geraads, D., Coppens, Y. & Guillemot, C. Taphonomical and paleoenvironmental study of OMO-33, a late pliocene hominid locality of the lower Omo Basin, Ethiopia. Rev. Paléobiol. 15, 339–347 (1996).
Boisserie, J. R. et al. New palaeoanthropological research in the Plio-Pleistocene Omo Group, lower Omo Valley, SNNPR (Southern Nations, nationalities and people Regions), Ethiopia. CR Palévol. 7, 429–439 (2008).
Suwa, G., White, T. D. & Howell, F. C. Mandibular postcanine dentition from the Shungura Formation, Ethiopia: Crown morphology, taxonomic allocations, and Plio-Pleistocene hominid evolution. Am. J. Phys. Anthropol. 101, 247–282 (1996).
Hlusko, L. J. et al. Taxonomic assignments for the 3.4 Ma to 1.1 Ma hominin postcanine teeth from the Usno Formation and the Shungura Formation, Lower Omo Valley, Ethiopia. Preprint at (2024). https://doi.org/10.31233/osf.io/g7kfx
Braun, D. R., Plummer, T. W., Ferraro, J., Ditchfield, P. & Bishop, L. Raw material quality and Oldowan hominin toolstone preferences: Evidence from Kanjera South, Kenya. J. Archaeol. Sci. 36, 1605–1614 (2009).
Braun, D. R. et al. Early Oldowan technology thrived during pliocene environmental change in the Turkana Basin, Kenya. Nat. Commun. 16, 9401 (2025).
Finestone, E. M. et al. Selective use of distant stone resources by the earliest Oldowan toolmakers. Sci. Adv. (2025).
Isaac, G. L. The activities of early African hominids: A review of archaeological evidence from the time Soan two and a half to one million years ago. in Human Origins: Louis Leakey and the East African Evidence (eds Isaac, G. L. & McCown, E. R.) 483–514 (Benjamin Inc., Philippines, 1976).
Toth, N. The artefact assemblages in the Ligh of experimental studies. in Koobi Fora Research Project. Volume 5: Plio-Pleistocene Archaeology (eds Isaac, G. L. & Isaac, B.) 363–401 (Oxford University Press, New-York, 1997).
Quade, J. et al. Paleoenvironments of the earliest stone toolmakers, Gona, Ethiopia. Geol. Soc. Am. Bull. 116, 1529–1544 (2004).
Reed, D. N. & Geraads, D. Evidence for a late pliocene faunal transition based on a new rodent assemblage from Oldowan locality Hadar AL 894, Afar Region, Ethiopia. J. Hum. Evol. 62, 328–337 (2012).
Plummer, T. W. et al. Oldest evidence of toolmaking hominins in a Grassland-Dominated ecosystem. PLoS ONE. 4, e7199 (2009).
Braun, D. R. et al. Early hominin diet included diverse terrestrial and aquatic animals 1.95 Ma in East Turkana, Kenya. PNAS 107, 10002–10007 (2010).
Mercader, J. et al. Earliest Olduvai hominins exploited unstable environments ~ 2 million years ago. Nat. Commun. 12, 1–15 (2021).
Renfrew, C. B.A.R., Oxford,. The Anatomy of Innovation. in Social Organization and Settlement vol. 47. 89–117 (1978).
Boisserie, J. R. et al. New paleoanthropological research in the Plio-Pleistocene Omo-Group, lower Omo Valley, SNNPR (Southern Nations, nationalities and people Regions), Ethiopia. C.R. Palevol. 7, 429–439 (2008).
Acknowledgements
This work was granted a permit by the Ethiopian Heritage Authority (EHA) as part of the Omo Group Research Expedition (OGRE). We thank the EHA curators for their unfailing help and logistical support during our study sessions at the National Museum of Ethiopia, as well as the staff members of the French Center for Ethiopian Studies (CFEE) in Addis Ababa, all field workers for their essential contribution, and the South Omo Zone, the Nyangatom and Dassanech Woredas and their people for their help and reception. We would like to thank James Clark for his highly relevant comments.
Funding
This study was supported by public grants from the French Ministry for Europe and Foreign Affairs (to J.-R.B.), from the French National Research Agency (ANR-project OLD, grant N° ANR-16-CE27-0009-01 to A.D.), and from the University of Bordeaux’s IdEx “Investments for the Future” program / GPR “Human Past” (to A.D).
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Anne Delagnes: Conceptualization, Field investigation, Methodology, Formal analysis, Writing, Data curation, Review and editing, Funding acquisition. Pascal Bertran: Conceptualization, Field investigation, Methodology, Writing, review and editing. Brad Gravina: Field investigation, Formal analysis, Writing, Review and editing. Frédéric Santos: Formal analysis, Methodology, Writing, Review and editing. Océane Spinelli Sanchez: Field investigation, Methodology, Formal analysis, Writing, Review and editing. Aline Galland: Field investigation, Writing, Data curation, Review and editing. Michel Brenet: Field investigation, Methodology, Formal analysis, Writing, Review and editing. Haregwin Belete Hailu: Field investigation, Formal analysis, Review and editing , Bisrat Gebreegziabher Araya: Field investigation, Formal analysis, Review and editing, Fikeru Mekonenn Sissay: Field investigation, Formal analysis, Review and editing, Misganaw Gebremichael Woldetsadik: Field investigation, Formal analysis, Review and editing, Jean-Renaud Boisserie: Field investigation, Formal analysis, Writing, Data curation, Review and editing, Funding acquisition.
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Delagnes, A., Bertran, P., Gravina, B. et al. A regional-scale mobility model for the early hominin occupation of the Lower Omo Valley (Ethiopia). Sci Rep (2026). https://doi.org/10.1038/s41598-026-39972-9
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DOI: https://doi.org/10.1038/s41598-026-39972-9


