Fig. 4: The frequency of different strategies in the cg − r plane in a structured population.

The time average frequencies of costly cooperators (a), costly defectors (b), non-costly cooperators (c), and non-costly defectors (d) in the cg − r plane are color plotted. The system shows a red queen dynamic in which cooperators only in the costly institution survive in large numbers (for smaller enhancement factors), or a black queen dynamic, where cooperators in both institutions survive and help each other to suppress defection (for larger enhancement factors). Parameter values: g = 5, nu = 10−3, and π0 = 2. The population resides on a 200 × 200 first nearest neighbor square lattice with von Neumann connectivity and periodic boundaries. The simulation is performed for 5000 time steps starting from an initial condition in which all the individuals are defectors and prefer one of the two institutions at random. The time average is taken over the last 2000 steps.