Fig. 5: Concept map of optimizing ley durations for continuous grasslands and crop rotations.

Carbon (C) maintained in the ecosystem is the balance between C input from plants and C output through heterotrophic respiration (Rh), which is mainly exhibited through the soil organic carbon (SOC) storage and partly through C in undecomposed residues (RC). a For continuous grasslands, adjusting the ploughing and renewal cycles of leys can maximize the benefit of SOC storage by balancing C input and Rh. b For crop rotations, adjusting the duration of leys can maximize the SOC storage without requiring long ley durations.