Fig. 1: Phylogeny and phylogeography of the genus Macrovipera.

The distribution ranges of the species M. lebetina (light brown), M. schweizeri (dark brown) and M. razii (dark blue), as well as the subspecies M. l. transmediterranea (of debated validity and possibly extinct), were sourced from the WHO Snakebite Information and Data Platform30. The distribution ranges of the other M. lebetina subspecies (ellipses) and the phylogenetic tree were adapted from Stümpel44, with the triangles at the branch tips representing the age and sampling of each taxon. The points on the map indicate the sampling localities from Stümpel’s work. The star represents M. lebetina populations from coastal southern Türkiye sharing mitochondrial haplotypes with M. schweizeri. Photo credits: Rami Khasab (M. l. obtusa), Seyed Mahdi Kazemi (M. razii), Ignazio Avella (M. schweizeri).