Table 1 Results of palynological analyses on the sample of the white powder from the heart of Richard I

From: The embalmed heart of Richard the Lionheart (1199 A.D.): a biological and anthropological analysis

Identified taxa (family, genus and species when possible)

Number of pollen grains

State of preservation of pollen grains

Period of pollination of referred plants

Present-day geographic distribution of referred plants

Pinus, Pinaceae

4

Poor

May-June

(Pinus silvestris (Scots pine), only one specie spontaneous in the north-western Massif Central

Quercus ilex (holm oak), Fagaceae

1

Moderate

April-May

Common in the Mediterranean region, but also distributed in Western France up to Brittany

Plantago (plantain), Plantaginaceae

1

Moderate

> April

Plantago species widely distributed in western France

Campanulaceae

1

Good

> April

Several genera (Campanula i.e. bellflower, Jasione, Phyteuma, etc.) widely distributed in France

Myrtus communis (myrtle), Myrtaceae

3

Good

May-July

Sweet-smelling plant endemic of the Mediterranean region

Asteraceae Asteroideae (chrysanthemums, maybe the species Leucanthemum vulgare (daisy) according to the small size of the pollen grains (<20 μm)

2

Good

May-August

Widely distributed in France

Lamiaceae (Mentha (mint), according to the ornamentation of exine)

1

Good

July-October

Mentha (mint) displays several species living in Western France such as M. silvestris (sweet-smelling plant; blooming July-September), M. viridis (August-October), M. aquatica (July-September), M. arvensis (strong-smelling plant: July-October) and M. Pulegium (strong-smelling plant: July-October)

Populus (poplar), Salicaceae

1

Good

March-April

Three species of Populus (P. alba, P. tremula, P. nigra) widely distributed in France