Figure 1

Major transformations of Mt1 in the evolution of avian opposable hallux.
(a) Basal dinosaur exhibit the Mt1 (grey arrow) articulated to distal tarsals (Herrerasaurus); (b) Mt1 of basal tetanurans had lost its proximal epiphysis and articulated to Mt2 (Allosaurus); (c) Basal birds had the Mt1 articulated to the distal Mt2 (Archaeopteryx); (d) Most extant birds exhibit the hallux opposed to other toes, as observed in late chicken embryo (HH44); (e) Detail showing hallux inverted orientation.