Table 3 Results of forward selection and Monte Carlo permutation tests from the redundancy analysis (RDA) of all 94 sampling sites.

Variables	Lambda-1	P	Lambda A	P
TN	0.05	0.002^**	0.05	0.002^**
Latitude	0.03	0.008^**	0.02	0.010^*
T	0.03	0.026^*	0.03	0.020^*
NO₃-N	0.02	0.176	0.02	0.152
Distance	0.01	0.556	0.01	0.370
Crustacea	0.01	0.888	0.00	0.500
NH₄-N	0.03	0.034^*	0.01	0.624
Longtitude	0.02	0.120	0.01	0.818
Protozoa	0.00	0.802	0.00	0.706
SD	0.01	0.546	0.01	0.784
TP	0.03	0.010^**	0.00	0.992
SRP	0.03	0.018^*	0.01	0.610
Altitude	0.01	0.178	0.00	1.000

Environmental variables were listed by the order of their inclusion in the RDA model. Lambda-1 represents the independent effect of each environmental variable when the variable was treated separately. Lambda A represents the effect that each variable brings in addition to the all variables already selected and the most important variable included first. The P-value indicates the significance of each variable either when considered independently or dependently (*P < 0.05, **P < 0.01). TN = total nitrogen; T = water temperature; NO₃-N = nitrate nitrogen; NH₄-N = ammonia nitrogen; distance = average geographical distance of a given sampling site from the other sampling sites; Crustacea = total abundance of all crustacean species; Protozoa = total abundance of all protozoa species; SD = Secchi disk depth; TP = total phosphorus; SRP = soluble reactive phosphorus.

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