Figure 3: Stepped movement of the CD.
From: Structural intermediates and directionality of the swiveling motion of Pyruvate Phosphate Dikinase

CDs of 5JVL/C (A, orange) and 5JVN (B, yellow). Helix 20 containing the catalytic H456 is depicted as cylinder with the Cα atom of H456 shown as red sphere. The rotational axis for the transitions between states A and B is depicted as a blue arrow. The distance between the Cα atoms of the catalytic His456 is shown as a dashed line.