Introduction

The meridional overturning circulation (MOC), connecting the deep ocean with the atmosphere, plays a central role in the climate system by transporting heat, fresh water and carbon across the globe. Model studies have consistently shown that the structure and strength of the MOC depend sensitively on the deep-ocean turbulent diapycnal mixing1,2,3,4,5,6 (hereafter referred to as mixing). The geographical distribution of mixing in the ocean is highly inhomogeneous7,8,9,10,11. Enhanced mixing is typically observed over rough topography, associated with the local breaking of internal tides (also known as baroclinic tides) generated by interactions of barotropic tides with rough topography12,13,14,15. The global energy conversion from barotropic tides to baroclinic tides is estimated to be approximately 1 TW16, a large proportion of the estimated power needed to maintain the MOC17.

Vertical structures of internal tides are usually described by the eigenmodes related to ocean stratification18. The modal partition of tidal energy conversion shapes the geographical pattern of mixing and is a cornerstone of tidal mixing parameterization3,19,20. On the one hand, low-mode (e.g., mode 1–3) internal tides, having small vertical shear and large horizontal group velocity, can radiate over a long distance away from their generation sites before dissipation, contributing to the background mixing over the global ocean21. On the other hand, high-mode (≥4) internal tides tend to dissipate locally due to their large vertical shear and slow horizontal group velocity, resulting in hotspots for mixing over rough topography12,22.

In most of the state-of-the-art coupled global climate models (CGCMs)23,24,25,26,27,28, the tidal energy conversion and its modal partition are simply assumed to be constant over time. However, the increased emission of greenhouse gases and subsequent ignition of a global warming has enhanced the stratification, especially, in the upper ocean29,30. There is a prevailing thought that the enhanced stratification would weaken ocean mixing by suppressing the instability processes responsible for turbulence generation31,32. However, this thought overlooks the important effects of stratification on the energy conversion from barotropic to baroclinic tides for different vertical modes33. There have been some regional studies showing that the enhanced stratification could result in locally enhanced or reduced tidal energy conversion34,35,36. Yet it remains unclear how the tidal energy conversion and its modal partition respond to global warming on a global scale. Such knowledge is necessary for a more consistent projection of future climate change, particularly in view of the crucial role of tidal mixing in maintaining the MOC1,2,3,4,5,6.

In this study, we investigate the response of tidal energy conversion and its modal partition to the enhanced stratification under global warming using a linear model of internal tide generation12 and simulations from an ensemble of global coupled climate models (CGCMs) in the Coupled Model Intercomparison Project phase 6 (CMIP6) under a high carbon emission scenario24 (‘CMIP6 CGCMs’ in Methods). We show that the changes in low-mode tidal energy conversion are spatially inhomogeneous, resulting in a nearly negligible increase (0.9 ± 0.2%) of the globally integrated conversion rate. In contrast, there is a universal increase of the high-mode tidal energy conversion, with its global integral projected to rise by 7.8 ± 0.7% by the end of this century, compared to its present level. This enhanced high-mode tidal energy conversion implies stronger deep-ocean tidal mixing over rough topography and needs to be properly parameterized in CGCMs for a more reliable projection of future climate changes.

Results

Projected future changes of ocean stratification under global warming

The tidal energy conversion and its modal partition depend on both the near-bottom (Nb) and depth-averaged buoyancy frequency (\(\bar{N}\)) (‘Computation of tidal energy conversion’ in Methods). On the one hand, the tidal energy conversion for any n-th mode En becomes larger with the increasing Nb. In particular, the dependence of En on Nb is the same among individual modes and almost linear provided that Nb is much larger than the tidal frequencies, which holds over many parts of the global ocean (Fig. 1a). On the other hand, the effect of \(\bar{N}\) on En is more complicated, depending on the spectral density of tidal energy conversion in the horizontal wavenumber space (‘Computation of tidal energy conversion’ in Methods). A larger (smaller) \(\bar{N}\) reduces (increases) the modal horizontal wavenumber Kn of internal tides for the n-th mode. For a negative slope of the spectral density of tidal energy conversion, a reduced Kn results in an increase of En. The opposite is true when the slope is positive.

Fig. 1: Response of near-bottom buoyancy frequency and depth-averaged buoyancy frequency to global warming.
figure 1

ad Geographical distribution of time-mean (1995–2004) near-bottom buoyancy frequency Nb and depth-averaged buoyancy frequency \(\overline{N}\) in the observation (a, b) and CMIP6 coupled global climate models (CGCMs) ensemble mean (c, d). e, f Projected changes (2091–2100 minus 1995–2004) of Nb and \(\overline{N}\) in the CMIP6 CGCM ensemble mean. Changes insignificant at a 95% confidence level are filled in white. Numbers in white represent the globally averaged changes in percentage as well as their 95% confidence interval. Areas shallower than 500 m are filled in gray and discarded in the analysis.

The climatological mean (1995–2004) Nb in the observations37 (‘Observational dataset’ in Methods) varies by an order of magnitude over the global ocean (Fig. 1a). The geographical distribution of Nb is tightly related to the sea floor depth, with larger values of Nb generally occurring in shallower regions with rougher topography (Supplementary Fig. 1; ‘Definition of sea floor roughness’ in Methods). There is also prominent geographical variability of climatological mean \(\bar{N}\) (Fig. 1b). In contrast to Nb, variability of \(\bar{N}\) is primarily controlled by the latitude, with large and small values residing in the tropical and polar oceans, respectively. Nevertheless, imprint of sea floor depth on the geographical distribution of \(\bar{N}\) is still noticeable in some regions (e.g., the mid-ocean ridges). The magnitudes and geographical distributions of climatological mean Nb and \(\bar{N}\) are well reproduced by the CMIP6 CGCM ensemble mean (Fig. 1c, d), lending support to these CGCMs’ fidelity in representing large-scale ocean stratification.

The value of \(\bar{N}\) in the CMIP6 CGCM ensemble mean exhibits a universal increase over the global ocean in response to global warming. Its global average during 2091–2100 is projected to be 10.2 ± 1.4% larger than that during 1995–2004 (Fig. 1f). In contrast, the response of Nb to global warming is weaker and heterogeneous (Fig. 1e), with patches of positive and negative trends. The globally averaged Nb is projected to increase only by 2.7 ± 0.5% from 1995–2004 to 2091–2100. It thus suggests that even under a high carbon emission scenario, ocean warming is mainly confined to the upper ocean by the end of 21st century and has a stronger impact on the global mean \(\bar{N}\) than Nb.

Projected future changes of tidal energy conversion and its modal partition under global warming

The tidal energy conversion into the first 50 modes E1–50 as well as its partition into low modes (E1–3) and high modes (E4–50) is estimated from the linear theory (‘Computation of tidal energy conversion’ in Methods; Supplementary Fig. 2). Here the computation is limited to the first 50 modes, as the resolution (15-arc second) of the bathymetric dataset (‘Observational dataset’ in Methods) does not allow to resolve higher modes38. Nevertheless, these unresolved higher modes have a much smaller energy conversion than the resolved ones39. The globally integrated climatological mean E1–50 is 864 GW. Most of this energy conversion occurs over the rough topography such as steep ridges and seamounts (Fig. 2a). The climatological mean E1–3 and E4–50 have similar geographical distributions to that of E1–50 (Fig. 2a–c). The globally integrated climatological mean E1–3 and E4–50 are 568 GW and 296 GW, respectively.

Fig. 2: Response of tidal energy conversion and its modal partition to global warming.
figure 2

ac, Geographical distribution of time-mean (1995–2004) of total tidal energy conversion E1–50 (a), and its partition into low modes E1–3 (b) and high modes E4–50 (c). df Same as ac, but for the projected changes (2091–2100 minus 1995–2004) under global warming. Changes insignificant at a 95% confidence level are filled in white. Numbers in white represent the globally integrated values as well as their 95% confidence interval.

Under global warming, E1–50 is increased over most parts of the global ocean (Fig. 2d). The increase of E1–50 is most evident over the rough topography where its climatological mean value is large. The increase of E1–50 is primarily attributed to E4–50. The globally integrated E1–50 increases by 28 ± 3.5 GW from 1995–2004 to 2091–2100, with E4–50 contributing to 23 ± 2.2 GW of this increase. In contrast, the change of E1–3 under global warming has both positive and negative values. The globally integrated E1–3 increases only by 5 ± 1.3 GW (Fig. 2e). Therefore, the effect of global warming on E1–50 is dominated by E4–50 whose global integral increases by about 8% by the end of 21st century, compared to its historical (1995–2004) level (Fig. 2c). We remark that such trends of tidal energy conversion and its modal partition do not only hold for the CMIP6 CGCM ensemble mean but also for most of the individual CGCM members (Fig. 3).

Fig. 3: Projected changes of globally integrated tidal energy conversion for individual CMIP6 coupled global climate models (CGCMs) members.
figure 3

Changes (2091–2100 minus 1995–2004) of total tidal energy conversion E1–50 (a), and its partition into low modes E1–3 (b) and high modes E4–50(c). Black dashed lines represent their ensemble mean results.

Both changes of Nb and \(\overline{N}\) can affect the response of tidal energy conversion and its modal partition to global warming. To separate their effects, the change of En under global warming is recomputed by either fixing Nb or \(\overline{N}\) at its historical (1995–2004) mean value. The changes of Nb and \(\overline{N}\) play a comparable role in the increase of E1–50 under global warming (Fig. 4a, d), contributing to 16 ± 5.1 GW and 12 ± 3.8 GW, respectively. However, they have distinct effects on the modal partition of tidal energy conversion (Fig. 4b, c, e, f). On the one hand, the enhanced Nb under global warming increases the global integrals of E1–3 and E4–50 by a similar fraction (2.1 ± 0.5% vs. 1.4 ± 0.7%). This is consistent with the nearly linear dependence of En on Nb (‘Computation of tidal energy conversion’ in Methods). On the other hand, the enhanced \(\overline{N}\) under global warming decreases the globally integrated E1–3 by 6 ± 0.6 GW, whereas it increases the globally integrated E4–50 by 18 ± 3.9 GW. As the effects of enhanced Nb and \(\overline{N}\) on E1–3 (E4–50) counteract (reinforce) each other, this explains the much larger increase of E4–50 than E1–3 under global warming.

Fig. 4: Effects of near-bottom buoyancy frequency and depth-averaged buoyancy frequency changes on the tidal energy conversion change under global warming.
figure 4

ac Geographical distribution of the projected changes (2091–2100 minus 1995–2004) of total tidal energy conversion E1–50 (a, d), and its partition into low modes E1–3 (b,e) and high modes E4–50 (c, f), caused by the changes of near-bottom buoyancy frequency Nb (ac) and depth-averaged buoyancy frequency \(\overline{N}\) (df), respectively. Changes insignificant at a 95% confidence level are filled in white. Numbers in white represent the globally integrated values as well as their 95% confidence interval.

The opposite effects of enhanced \(\overline{N}\) on E1–3 and E4–50 under global warming can be understood based on the reduced Kn in response to the enhanced \(\overline{N}\). This is demonstrated by using the M2 internal tides as a representative example (Supplementary Fig. 3). The energy conversion into the M2 internal tides is the main contributor to the total tidal energy conversion40 and its change under global warming (Fig. 2 and Supplementary Fig. 4). Specifically, the geographical distribution of time-mean \({E}_{{{{\rm{M}}}}_{2}}^{1-50}\) during 1995–2004 is highly consistent with that of E1–50 with a correlation coefficient of 0.97 (Fig. 2a and Supplementary Fig. 4a). Under global warming, \({E}_{{{{\rm{M}}}}_{2}}^{1-50}\) shows an increase over the major part of the global ocean with its globally integrated increase dominated by that of \({E}_{{{{\rm{M}}}}_{2}}^{4-50}\) (Supplementary Fig. 4).

The spectral density \({\varPsi }_{{{{\rm{M}}}}_{2}}(K)\) of M2 tidal energy conversion in the horizontal wavenumber space, tightly related to the topographic spectrum, shows distinct features between the regions with enhanced and reduced \({E}_{{M}_{2}}^{1-3}\) under global warming. In regions where \({E}_{{M}_{2}}^{1-3}\) is enhanced, the topographic spectrum rolls off rapidly (Supplementary Fig. 5a) and \({\varPsi }_{{{{\rm{M}}}}_{2}}(K)\) decreases monotonically as K increases. (Fig. 5a). As \({K}_{{M}_{2}}^{n}\) is reduced in response to the enhanced \(\overline{N}\) (Supplementary Fig. 3), this leads to larger \({\varPsi }_{{{{\rm{M}}}}_{2}}({K}_{{M}_{2}}^{n})\) for any n under global warming, increasing both \({E}_{{M}_{2}}^{1-3}\) and \({E}_{{M}_{2}}^{4-50}\). In regions with reduced \({E}_{{M}_{2}}^{1-3}\), the topographic spectrum becomes relatively flat (Supplementary Fig. 5b) and \({\varPsi }_{{{{\rm{M}}}}_{2}}(K)\) peaks around \({K}_{{M}_{2}}^{3}\) and attenuates towards both sides (Fig. 5b). Consequently, the reduced \({K}_{{M}_{2}}^{n}\) under global warming leads to an increase (decrease) of \({E}_{{M}_{2}}^{4-50}\) (\({E}_{{M}_{2}}^{1-3}\)) there. In summary, the reduced \({K}_{{M}_{2}}^{n}\) always increases \({E}_{{M}_{2}}^{4-50}\), whereas its effect on \({E}_{{M}_{2}}^{1-3}\) is region-dependent and relies on the shape of topography spectrum. This explains why the enhanced \(\overline{N}\) under global warming leads to an increase in E4–50 but an overall decrease in E1–3.

Fig. 5: Effects of modal horizontal wavenumber change on the energy conversion to M2 internal tides.
figure 5

Spectral density of M2 tidal energy conversion (\({\varPsi }_{{{\rm M}}_{2}}\)) as a function of horizontal wavenumber normalized by the mode-1 horizontal wavenumber of M2 internal tides \({K}_{{M}_{2}}^{1}\) during 1995–2004, averaged over the regions with the increased (a) and decreased (b) \({E}_{{{{\rm{M}}}}_{2}}^{1-3}\) under global warming. The color shadings denote the interquartile ranges of \({K}_{{M}_{2}}^{2}/{K}_{{M}_{2}}^{1}\), \({K}_{{M}_{2}}^{3}/{K}_{{M}_{2}}^{1}\) and \({K}_{{M}_{2}}^{4}/{K}_{{M}_{2}}^{1}\).

Discussion

In this study, the response of tidal energy conversion and its modal partition to global warming is evaluated by applying a linear model of internal tide generation to an ensemble of CGCM simulations under a high carbon emission scenario. Although E1–3 remains nearly unchanged, E4–50 is projected to rise by about 8% by the end of the 21st century, compared to its historical (1995–2004) level. It should be noted that the enhanced E4–50 and its dominant contribution to the increase of E1–50 is a qualitatively robust response to global warming, regardless of the time periods (Supplementary Fig. 6) and carbon emission scenarios (Supplementary Fig. 7). In particular, E4–50 is projected to rise by about 5% by the end of the 21st century under a medium carbon emission scenario41 that may be accepted by most countries pursuing sustainable growth.

Both the intensified near-bottom stratification and depth-averaged stratification play an important role in the changes of tidal energy conversion under global warming. However, their effects on the modal partition of tidal energy conversion are distinct from each other. The intensified near-bottom stratification increases both E1–3 and E4–50, whereas the intensified depth-averaged stratification increases E4–50 but reduces E1–3.

Our results suggest that the enhanced stratification under global warming increases energy conversion into high-mode internal tides which largely break near their generation sites and power the deep-ocean mixing locally7,12. Such an effect alone would enhance the deep-ocean mixing over the rough topography, leading to “hotter” mixing hotspots in a warming climate. However, the enhanced stratification also acts to suppress mixing as indicated by the Osborn relation42. Here we use a tidal mixing parameterization for high-mode internal tides43 (‘Tidal mixing parameterization for high-mode internal tides’ in Methods) to explore the relative importance of these two counteracting effects of enhanced stratification on deep-ocean mixing.

Figure 6 shows the parameterized deep-ocean (bottom 1000 m) diapycnal diffusivity κρ resulting from the breaking of high-mode internal tides in the historical period and its future change under global warming. The magnitude of historical κρ varies substantially in space, with mixing hotspots (κρ in the order of O(10−3 m2s−1)) over rough topography (Fig. 6a). Under global warming, the value of κρ increases over most parts of the global ocean, with the mixing hotspots generally becoming “hotter” (Fig. 6b). It thus suggests that the effect of enhanced stratification on energy conversion into high-mode internal tides plays a dominant role in the response of κρ to global warming. The magnitude of globally averaged κρ during 2091–2100 is projected to increase by 6% compared to its historical level. But locally this value can reach more than 20%.

Fig. 6: Implications of enhanced energy conversion into high-mode internal tides under global warming on the deep-ocean mixing.
figure 6

a Geographical distribution of vertical mean (bottom 1000-m) parameterized diapycnal diffusivity κρ caused by the breaking of high-mode internal tides during 1995–2004. b Same as a but for the difference between 1995–2004 and 2091–2100. Changes insignificant at a 95% confidence level are filled in white.

The increase of κρ caused by enhanced E4–50 may have important implications on the MOC and dianeutral upwelling4,44,45. Our preliminary analyzes suggest that it could accelerate the lower limb of the Atlantic MOC (AMOC) by about 10% (Supplementary Note 1 and Fig. 8). This change is comparable in magnitude to the projected slowdown (~10%) of the lower limb of the AMOC under a high carbon emission scenario caused by the intrusion of the Antarctic Bottom Water46. Furthermore, there is a significant response of the dianeutral upwelling to the enhanced E4–50 (Supplementary Note 2 and Fig. 9). The peak of the globally integrated dianeutral upwelling increases by about 10%. In view of the uncertainties in the tidal mixing parameterization42, the above results should be treated as suggestive rather than definitive. Nevertheless, they suggest important effects of the enhanced E4–50 resulting from the future stratification changes on tracer and mass transports in the deep ocean. So far the interactivity of parameterized tidal mixing is either neglected or limited in the state-of-the-art CGCMs23,24,25,26,27,28. Integrating stratification changes into tidal mixing parameterizations is imperative for accurately projecting anthropogenic climate changes in the future.

Methods

Observational dataset

We calculate the historical N using the climatological mean (1995–2004) temperature and salinity fields on a 0.25° × 0.25° grid from the World Ocean Atlas 2018 (WOA18)37. There are 102 vertical levels in the ocean with a maximum grid size of 100 m at the 5500 m depth. The two-dimensional topographic spectrum is computed from the updated Shuttle Radar Topography Mission dataset (SRTM15+)47 with a nominal resolution of 15-arc second.

CMIP6 CGCMs

In this study, a total of 25 CGCMs from the CMIP624 (Supplementary Table 1) are used to estimate the future change of N under the Shared Socioeconomic Pathways (SSP585), denoted as ΔN. The value of ΔN is computed as the simulated time-mean N during 2091–2100 minus that during 1995–2004. To remove the model drift, the counterpart of N change (the last decade minus the decade 96 years ago) in the pre-industrial control (PI-CTRL) simulation is subtracted from ΔN. When computing the tidal energy conversion during 2091–2100, we add ΔN to the observational time-mean N during 1995–2004 to minimize the model bias in the simulated N during the historical period.

Computation of tidal energy conversion

The tidal energy conversion rate is estimated using St. Laurent and Garrett’s12 formulation based on the linear theory proposed by Bell48,49:

$$\psi (K,\theta )=\frac{1}{2}{\rho }_{0}\frac{{[({N}_{b}^{2}-{\omega }^{2})({\omega }^{2}-{f}^{2})]}^{1/2}}{\omega }\times ({u}_{e}^{2}{\cos }^{2}\theta+{v}_{e}^{2}{\sin }^{2}\theta )K \phi (K,\theta )$$
(1)

where ψ(K, θ) is the spectral density of tidal energy conversion rate in the horizontal wavenumber space, ρ0 is the reference density, Nb is the near-bottom buoyancy frequency computed as the vertical mean buoyancy frequency within 0–300 m above the sea floor, \(\omega\) is the tidal frequency, f is the Coriolis frequency, ue (ve) is the semi-major (semi-minor) component of the barotropic tidal velocity amplitude retrieved from TPXO850, \(K={({k}^{2}+{l}^{2})}^{1/2}\) is the total horizontal wavenumber with k and l the respective wavenumbers along the semi-major and semi-minor directions, θ = arctan(l/k) is the azimuthal angle, and ϕ is the two-dimensional topographic spectrum. The azimuthally averaged spectral density of tidal energy conversion rate can be computed as:

$$\varPsi (K)=\frac{1}{2\pi }{\int }_{0}^{2\pi }\psi (K,\theta )Kd\theta$$
(2)

The vertical modes of internal tides can be determined by solving the Sturm-Liouville problem:

$$\frac{{d}^{2}{a}_{n}}{d{z}^{2}}+{c}_{n}^{-2}{N}^{2}{a}_{n}=0$$
(3)

with homogeneous boundary conditions at the sea floor (z = −H) and the sea surface (z = 0)

$${a}_{n}(0)={a}_{n}(-H)=0$$
(4)

where an and cn are the eigenfunction and eigenspeed for the n-th mode. Once cn is obtained, the modal horizonal wavenumber can be calculated based on the dispersion relation of internal waves as:

$${K}^{n}=\frac{\sqrt{{\omega }^{2}-{f}^{2}}}{{c}_{n}}$$
(5)

Finally, the tidal energy conversion rate into the n-th mode is given by:

$${E}^{n}={\int }_{{K}^{n}-\delta K/2}^{{K}^{n}+\delta K/2}\varPsi (K)KdK$$
(6)

where \(\delta K={K}^{n+1}-{K}^{n}\).

It should be noted that Eq. (1) derived from the linear theory is only valid for subcritical topography, i.e.,

$$\gamma=\frac{|\nabla h|}{{[({\omega }^{2}-{f}^{2})/({N}_{b}^{2}-{\omega }^{2})]}^{1/2}} < 1$$
(7)

where |h| and \({[({\omega }^{2}-{f}^{2})/({N}_{b}^{2}-{\omega }^{2})]}^{1/2}\) are the slopes of topography and radiated internal tide beams, respectively. The condition for subcritical topography is satisfied over most parts of the ocean, except for shallow regions with strong Nb. For this reason, the computation of Eq. (6) is confined to regions with water depth greater than 500 m. This neglects the downward radiation of internal tides generated in the regions shallower than 500 m to the deep ocean, which is suggested to contribute little to the deep ocean energy budget of internal tides20. For areas with supercritical topography (γ > 1), a correction is implemented on En by dividing En by γ2, following Green and Nycander51 and Vic et al.38.

The values of En are computed for the three main tidal constituents (M2, S2, and K1), which collectively account for about 90% of the total tidal energy conversion52. Following de Lavergne et al.19, we apply a scaling factor to estimate the total energy conversion for the eight principal tidal constituents (M2, S2, N2, K2, K1, O1, P1, Q1):

$${E}^{n}=1.05\times {E}_{{M}_{2}}^{n}+1.09\times {E}_{{{{\rm{S}}}}_{2}}^{n}+1.70\times {E}_{{{{\rm{K}}}}_{1}}^{n}$$
(8)

Definition of sea floor roughness

The sea floor roughness is calculated as the root-mean-square of topography height in each 0.5 × 0.5° grid cell. Before computing the sea floor roughness, the large-scale topography is removed by subtracting a fitted polynomial plane of topography height53.

Tidal mixing parameterization for high-mode internal tides

The mixing driven by the local dissipation of high-mode internal tides is parameterized as38,43:

$${\kappa }_{\rho }=\frac{\varGamma {E}^{4-50}F(z)}{{\rho }_{0}{N}^{2}}$$
(9)

where Γ = 0.2 is the mixing efficiency42 and F(z) is the vertical structure function defined as:

$$F(z)=\frac{{e}^{(z-H)/\eta }}{\eta (1-{e}^{-H/\eta })}$$
(10)

with a vertical decay scale of η = 500 m.