Fig. 1: Map of the study area.

a, The location of Sulawesi within the Wallacean archipelago (Wallacea)—the zone of oceanic islands between the Asian and Australian continental regions (Sunda and Sahul, respectively). MSL, mean sea level. b, The southwestern peninsula of Sulawesi, showing the locations of the Talepu (1) and Calio (2) sites on opposite sides of the Walanae River. Sources of map data: General bathymetric maps (https://www.gebco.net/data_and_products/gridded_bathymetry_data/); DEMNAS Badan Informasi Geospasial (https://tanahair.indonesia.go.id/demnas/#/demnas).