Figure 3

Ex vivo crypt-villous organoid cultures. (a) A representative mouse crypt cultured ex vivo in gelmatrix that grew into a crypt-villous organoid with budding crypts (black arrows) extending from the surface at days 1–11. (b) Organoid illustration showing lumen, villus domain, crypt domain and ISCs in green (from Sato et al28). (c) Exponential growth of crypt-villous organoids cultured with R-spondin 1, Noggin and HB-EGF (50 ng/ml).