Table 1 Principal component loadings from the correlation analysis.

Test	Behavioral parameter	Biological meaning of the metric	Principal component loadings^a
Test	Behavioral parameter	Biological meaning of the metric	PC1 Exploration	PC2 Boldness	PC3 Locomotor/activity	PC4
JBT	JBS	Decision-making under ambiguity	0.131	0.717	0.081	−0.179
OFT	Mean distance moved	Mobility	−0.571	−0.009	−0.645	0.374
	Time spent in the inner zone	Spatial preference	0.906	0.264	−0.031	0.017
	Number of crossings between the inner and outer zones	Spatial examination	0.780	0.441	−0.282	0.114
	Mean distance from walls	Edge-orienting	0.885	0.421	−0.050	0.007
	Mean angular velocity	Mobility	−0.804	0.139	0.200	−0.370
NOT	Mean distance moved	Mobility	−0.491	0.409	−0.651	−0.157
	Latency to the first approach to the novel object	Willingness to take risks	−0.143	0.707	−0.209	−0.162
	Time spent in the novel object zone	Object examination	0.261	−0.594	−0.517	−0.486
	Number of crossings between the novel object and empty zones	Object examination	0.496	−0.672	−0.346	−0.319
	Mean angular velocity	Mobility	−0.280	0.548	−0.117	−0.598
		Eigenvalue^b	3.861	2.750	1.417	1.071
		% Variance explained	35.107	25.007	12.885	9.741

Loadings extracted by principal component analysis from the correlation matrix of behaviors across tests for zebrafish belonging to the upper and lower quartiles of the JBS. Bold type indicates the strongest contributors (coefficient > 0.64) to each principal component (PC).
^aCorrelation between principal components and variable values.
^bVariance of transformed data used for each component.

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