Table 2 Structure of the river network according to suitability and overlap with PAs within each ecological cluster and on the interconnecting paths; the contribution of each variable to the overall ecological cost, computed as percentage of the total number of raster cells on the cluster’s interconnecting network for which the variable cost was positive (FFP—flash flood potential; RWQ—remote water quality; ALT—altitude; MMT—multiannual mean temperature).

Cluster	% suitable	% PA	Mean cost	% cost per variable
Cluster	% suitable	% PA	Mean cost	FFP	RWQ	ALT	MMT
1	84.9	61.22	0.061	0	100	0	0
2	53.17	39.24	0.142	0.04	93.77	4.46	6.15
3	87.57	91.52	0.048	0	100	0	0
4	40.8	26.11	0.127	0	74.51	13.9	24.84
5	39.56	55.6	0.141	0.3	58.37	38.23	40.73
6	37.75	19.6	0.108	0.54	38.07	56.42	62.27
7	82.58	100	0.033	0	100	0	0
8	83.1	100	0.026	0	77.90	14.36	22.1
9	95.51	25	0.009	0	60.71	14.29	39.29
10	61.47	38.68	0.072	0	70.38	23.17	29.35
11	36.48	37.15	0.161	0	94.65	63.05	63.58
12	61.85	51.69	0.073	1.31	59.68	55.11	59.48
13	55.78	81.66	0.151	0	80.06	96.04	95.96
14	64.54	48.35	0.068	0	69.85	17.21	28.08
15	36.9	24.34	0.173	0	88.13	62.86	77.45
All	52.27	43.57	0.110	0.23	72.08	37.09	43.51
Connect	3.23	51.12	0.383	0	96.86	84.78	66.21

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