Table 2 Summary of the variation of meristic characters and measurements (mm) in Elaphe druzei sp. nov. (own data), E. sauromates and E. urartica (Jablonski et al.25 and Jablonski pers. data).

From: A new, rare, small-ranged, and endangered mountain snake of the genus Elaphe from the Southern Levant

Meristic characters

Elaphe druzei sp. nov

E. sauromates

E. urartica

(n = 22)

(n = 63)

(n = 32)

Preoculare

1–2

1–3

1–3

(1.35 ± 0.49)

(1.89 ± 0.52)

(1.52 ± 0.59)

Postoculare

2–3

1–3

1–3

(2.06 ± 0.24)

(2 ± 0.17)

(2.03 ± 0.25)

Temporale

2–4

1–3

2–3

(2.35 ± 0.49)

(2 ± 0.17)

(2.46 ± 0.63)

Posttemporale

2–4

2–5

2–4

(3.47 ± 0.71)

(3.40 ± 1.02)

(3.58 ± 0.61)

Supralabialia

8–9

7–10

8–9

(8.06 ± 0.24)

(8.16 ± 0.42)

(8.02 ± 0.13)

Supralabialia contacting the eye

4th and 5th (n = 12)

4th and 5th (n = 34)

4th and 5th (n = 13)

4th and 5th, and 5th and 6th (n = 2)

4th and 5th, and 5th and 6th (n = 4)

4th and 5th, and 5th and 6th (n = 1)

 

5th and 6th (n = 1)

 
 

5th and 6th, and 6th and 7th (n = 1)

 
 

5th and 6th, and 4th, 5th and 6th (n = 1)

 

Sublabialia

10–12

9–12

9–13

(10.71 ± 0.59)

(10.72 ± 0.73)

(10.90 ± 0.72)

Gulars in a transverse row between the last two sublabialia

11–17

10–16

11–16

(14.59 ± 1.54)

(13.78 ± 1.31)

(14.23 ± 1.34)

Gulars touching anterior inframaxillaries

Absent or 1–2

Absent or 1–2

1–2

(1.24 ± 0.56)

(1.06 ± 0.35)

(1.59 ± 0.50)

Gulars between posterior inframaxillaries

Absent or 1–4

Absent or 1–4

2–5

(2.53 ± 1.07)

(2.52 ± 0.83)

(2.45 ± 0.80)

Dorsalia and temporalia touching parietals

9–14

9–20

9–15

(11.37 ± 1.67)

(13.11 ± 2.11)

(11.32 ± 1.49)

Preventrale

2–6

Absent or 1–3

Absent or 1–3

(3.71 ± 1.38)

(1.41 ± 0.83)

(1.23 ± 0.61)

Ventralia

200–222

194–222

190*–211 (201.81 ± 5.29)

(208.60 ± 5.95)

(206.57 ± 5.84)

(199.64 ± 11.42)

Dorsalia

211–230

NA

NA

(218.00 ± 6.36)

  

Dorsalia at forebody

23 (n = 2), 25 (n = 15)

21 (n = 2), 22 (n = 2), 23 (n = 5), 24 (n = 4), 25 (n = 12), 27 (n = 1)

23 (n = 1), 24 (n = 2), 25 (n = 19)

Dorsalia at midbody

21 (n = 2), 23 (n = 8), 25 (n = 10)

23 (n = 6), 24 (n = 4), 25 (n = 50), 27 (n = 1)

23 (n = 4), 24 (n = 2), 25 (n = 26)

Dorsalia at hind body

19 (n = 15), 21 (n = 2)

18 (n = 1), 19 (n = 19), 20 (n = 4), 21 (n = 1)

18 (n = 1), 19 (n = 21)

Subcaudalia

57–78

61–79

60–74

(66.81 ± 5.33)

(70.84 ± 5.29)

(67.31 ± 4.08)

Anale

2

2

2

Measurements (in mm)

Elaphe druzei sp. nov

E. sauromates

E. urartica

(n = 22)

(n = 63)

(n = 20)

Snout-vent length

325–1541

278–1250

260–970

(1066.80 ± 376.22)

(778.58 ± 289.31)

(683.66 ± 216.49)

Tail length

52–294

54–323

50–245

(189.81 ± 81.34)

(175.66 ± 74.97)

(158.23 ± 60.47)

Total length

377–1835

334–1503

310–1205

(1228.29 ± 459.17)

(937.25 ± 351.87)

(841.88 ± 274.17)

Rostrum height

2.19–6.31

2.05–6.90

2.80–6.08

(4.56 ± 1.17)

(4.76 ± 1.45)

(4.06 ± 0.75)

Rostrum width

3.43–9.48

3.10–10.24

4.00–8.02

(7.14 ± 1.71)

(6.71 ± 1.94)

(6.04 ± 1.49)

Inter-nostril distance

4.51–10.79

2.40–10.54

4.30–8.88

(8.72 ± 2.06)

(7.21 ± 2.20)

(6.23 ± 1.56)

Loreal length

2.14–4.63

NA

NA

(3.29 ± 0.80)

  

Eye diameter

3.51–7.49

3.40–6.54

3.35–6.62

(5.55 ± 1.18)

(4.99 ± 1.01)

(4.42 ± 0.85)

Head length

15.00–44.00

14.30–38.70

16.85–31.49

(30.09 ± 7.56)

(25.02 ± 6.44)

(23.86 ± 5.81)

Head width

7.53–24.00

10.90–22.80

16.64–21.53

(15.71 ± 4.70)

(19.13 ± 3.91)

(19.62 ± 1.74)

Head height

7.15–21.83

NA

NA

(14.67 ± 3.83)

  

Supraoculare width

2.44–5.99

2.00–6.90

2.80–5.08

(4.72 ± 1.109)

(4.31 ± 1.39)

(3.81 ± 0.73)

Frontale length

3.94–9.51

4.30–12.50

4.80–8.48

(6.76 ± 1.45)

(7.98 ± 1.91)

(6.67 ± 1.31)

Frontale width

4.79–8.09

3.50–8.60

3.5–6.7

(8.00 ± 1.51)

(5.91 ± 1.45)

(5.27 ± 1.15)

Anterior inframaxillaries length

5.32–14.24

4.50–14.06

6.70–9.70

(10.32 ± 2.74)

(8.43 ± 2.70)

(7.85 ± 0.85)

Posterior inframaxillaries length

3.84–10.96

3.40–12.50

5.20–8.45

(7.75 ± 2.14)

(7.41 ± 2.49)

(6.38 ± 1.15)

  1. Data presented as minimum–maximum (average ± standard error); n = number of examined specimens.
  2. *Referring to the paratype of E. urartica from Azerbaijan (AZ IZANAS 518) that was incorrectly stated as having 154 ventral scales in Jablonski et al.25 and the number was corrected by Bunyatova, in lett. (2022).