Table 2 Species composition classified by order and family among seasons and the biomass (unit: g) and abundance (unit: ind.) of each species in the artificial oyster reef areas and control area. The symbol ● means present in this month and season. Values in parentheses represent the niche width index.—means no records. B and N represent the biomass and number.

Aciculata

 Aphroditidae

  Aphrodita talpa

–

●(0.38)

●

●(0.09)

●(0.33)

●

11

Acropomatiformes

 Lateolabracidae

  Lateolabrax japonicus

Japanese seabass

●(0.03)

●

1

Amphilepidida

 Amphiuridae

  Amphiura vadicola

–

●(0.06)

●

1

Clupeiformes

 Clupeidae

  Konosirus punctatus

Dotted gizzard shad

●(0.03)

●

1

 Engraulidae

  Thryssa kammalensis

Thryssa kammalensis

●(0.25)

●(0.04)

●

24

Decapoda

 Alpheidae

  Alpheus distinguendus

Forceps snapping shrimp

●(0.57)

●

●(0.06)

●

●(0.12)

8

4

  Alpheus japonicus

Japanese snapping shrimp

●(0.24)

●(0.07)

●

●(0.11)

●

●(0.08)

●(0.14)

●

●(0.63)

8

28

 Crangonidae

  Crangon affinis

Hakodate sand shrimp

●(0.44)

●(0.11)

●

●(1.96)

●

●(0.02)

105

1

 Diogenidae

  Diogenes edwardsii

Edward’s hermit crab

●(0.37)

●

●(0.03)

●

●(1.89)

17

153

 Dorippidae

  Dorippe japonica

–

●(0.09)

●

●(0.24)

●

●(0.60)

5

22

 Goneplacidae

  Eucrate alcocki

–

●(0.07)

●

1

 Leucosiidae

  Philyra platycheir

–

●(0.47)

20

 Lysmatidae

  Lysmata vittata

Indian lined shrimp

●(0.96)

●

●(0.12)

●

●(0.11)

●(0.03)

156

1

 Majidae

  Hyastenus diacanthus

Horn decorator crab

●(0.30)

●

4

  Pugettia nipponensis

–

●(0.34)

●(0.05)

●

●(0.37)

●

27

 Palaemonidae

  Palaemon gravieri

Chinese ditch prawn

●(1.25)

●(0.45)

●

●(0.21)

●(0.10)

●

●(0.40)

●

●(0.84)

●(2.49)

●

●(2.55)

153

353

 Pasiphaeidae

  Leptochela gracilis

Lesser glass shrimp

●(0.07)

●(0.07)

●

●(0.43)

●

●(0.40)

39

30

 Pinnotheridae

  Xenophthalmus pinnotheroides

Blind pea crab

●(0.39)

●

6

 Portunidae

  Charybdis japonica

Japanese swimming crab

●(0.18)

●(1.57)

●

●(1.70)

●(1.34)

●

●(2.79)

●

●(1.08)

●(0.14)

●

●(2.46)

1180

210

  Portunus trituberculatus

Gazami crab

●(0.11)

●

●(0.28)

1

5

 Sergestidae

  Acetes chinensis

Northern mauxia shrimp

●(0.11)

●

●(0.07)

3

2

Forcipulatida

 Asteriidae

  Asterias amurensis

North Pacific seastar

●(0.40)

●(1.09)

●

●(0.08)

●

●(0.22)

●(1.44)

●

●(0.08)

246

3

Gastropoda

 Naticidae

  Neverita didyma

Bladder moon snail

●(0.03)

●(0.03)

●

●(0.15)

●

●(0.06)

5

2

Gobiiformes

 Gobiidae

  Chaeturichthys stigmatias

Branded goby

●(0.13)

●

●(0.80)

●

●(0.11)

●

●(2.35)

27

190

  Synechogobius ommaturus

Asian freshwater goby

●(1.45)

●

●(1.48)

●(1.59)

●

●(2.68)

●

●(1.24)

●(3.48)

●

●(0.66)

773

46

  Tridentiger barbatus

Shokihaze goby

●(0.52)

●(0.13)

●

●(1.27)

●

●(0.19)

31

8

  Tridentiger bifasciatus

Shimofuri goby

●(0.15)

●

3

  Tridentiger trigonocephalus

Chameleon goby

●(0.05)

●

1

Haplosclerida

 Chalinidae

  Haliclona similis

–

●(0.12)

●

6

Myopsida

 Loliginidae

  Loliolus japonica

Japanese squid

●(0.08)

●

4

Nuculida

 Nuculidae

  Ennucula tenuis

Smooth nutclam

●(0.07)

●

2

Neogastropoda

 Muricidae

  Rapana venosa

Purple whelk

●(0.06)

●(0.87)

●

●(0.46)

●(0.32)

●

●(0.43)

●

●(0.30)

99

10

 Nassariidae

  Nassarius siquijorensis

–

●(0.08)

●

●(0.54)

●

14

  Nassarius variciferus

–

●(0.10)

●

●(0.02)

2

1

Octopoda

 Octopodidae

  Octopus variabilis

Whiparm octopus

●(0.16)

●(0.12)

●

●(0.06)

●(0.08)

●

●(0.13)

●

14

  Amphioctopus fangsiao

Gold-spot octopus

●(0.46)

●

●(0.15)

●

14

Perciformes

 Cottidae

  Trachidermus fasciatus

Roughskin sculpin

●(0.54)

●

●(0.03)

6

1

 Hexagrammidae

  Hexagrammos agrammus

Spotty-bellied greenling

●(0.09)

●

●(0.10)

●

5

  Hexagrammos otakii

Fat greenling

●(1.58)

●(0.32)

●

●(0.35)

●(0.03)

●

●(0.37)

●

●(0.74)

●(0.43)

●

●(0.09)

126

3

 Pholidae

  Pholis fangi

–

●(0.18)

●

●(0.08)

●

●(1.05)

●

●(0.41)

●

●(0.19)

47

7

 Platycephalidae

  Platycephalus indicus

Bartail flathead

●(0.04)

●

●(0.08)

●(0.03)

●

4

 Sciaenidae

  Johnius belangerii

Belanger’s croaker

●(0.76)

30

 Sebastidae

  Sebastes schlegelii

Korean rockfish

●(1.37)

●(1.21)

●

●(1.44)

●(0.50)

●

●(0.75)

●

●(1.40)

●(1.82)

●

●(0.90)

636

37

 Sillaginidae

  Sillago sihama

Silver sillago

●(0.03)

●

1

 Sparidae

  Acanthopagrus schlegelii

Blackhead seabream

●(0.03)

●

●(0.13)

●

4

 Stichaeidae

  Chirolophis japonicus

–

●(0.11)

●

2

  Ernogrammus hexagrammus

Six-lined prickleback

●(0.06)

●

1

 Zoarcidae

  Zoarces elongatus

–

●(0.09)

●

1

Pleuronectiformes

 Cynoglossidae

  Cynoglossus joyneri

Red tonguesole

●(0.05)

●

●(0.10)

●(0.08)

●

●(0.06)

6

2

  Cynoglossus robutus

–

●(0.02)

●

1

 Paralichthyidae

  Paralichthys olivaceus

Bastard halibut

●(0.33)

●

●(0.03)

10

1

 Pleuronectidae

  Platichthys bicoloratus

Stone flounder

●(0.23)

●(0.03)

●

●(0.24)

●

8

Semaeostomeae

 Ulmaridae

  Aurelia aurita

Moon jelly

●(0.09)

●

8

Stomatopoda

 Squillidae

  Oratosquilla oratoria

Japanese squillid mantis shrimp

●(0.17)

●

●(0.25)

●(0.08)

●

●(0.27)

●

●(0.18)

●

●(0.19)

29

4

Temnopleuroida

 Temnopleuridae

  Temnopleurus hardwickii

Hardwick’s sea urchin

●(0.04)

●

●(0.08)

●

●(0.07)

●

5

Tetraodontiformes

 Tetraodontidae

  Takifugu niphobles

–

●(0.09)

●

●(0.03)

●

7