Abstract
Current approaches to investigate the role of neural oscillations in natural scene processing have been limited to artificial stimuli and long data collection. We present a new way to decode real-world scenes participants are viewing from the steady-state visually evoked potentials (SSVEPs) evoked while wearing flickering LCD glasses. We discovered that SSVEP responses from real world scenes are surprisingly complex and have distinct waveform shapes: they differ markedly across scenes and participants but are consistent within individuals, even across multiple days. SSVEP shape varies greatly between stimuli, but is reliable, meaning that decoding works even with a single electrode. Decoding is highly accurate with 5–10 s of data and was still above chance level with less than a second of data. Decomposing the SSVEPs into frequency bands showed that the information about the visual scene is present across all of the harmonics of the flicker frequency: optimal decoding used the broadband signals, but with 40 Hz (gamma band) showing the highest amount of information after band-pass filters. These findings implicate a broad range of oscillations in encoding real-world scenes, with a particular importance for 40 Hz. The SSVEP’s temporal profile is a rich source of information for decoding.
Data availability
All raw data and example code are available from the Open Science Framework project page: https://doi.org/10.17605/OSF.IO/NF6VM.
References
Ladouce, S., Donaldson, D. I., Dudchenko, P. A. & Ietswaart, M. Understanding minds in real-world environments: toward a mobile cognition approach. Front. Hum. Neurosci. https://doi.org/10.3389/fnhum.2016.00694 (2017).
Vigliocco, G. et al. Ecological brain: Reframing the study of human behaviour and cognition. Royal Soc. Open. Sci. 11 (11), 240762. https://doi.org/10.1098/rsos.240762 (2024).
Gramann, K., Jung, T. P., Ferris, D., Lin, C. T. & Makeig, S. Toward a new cognitive neuroscience: Modeling natural brain dynamics. Front. Hum. Neurosci. 8 https://doi.org/10.3389/fnhum.2014.00444 (2014).
Shamay-Tsoory, S. G. & Mendelsohn, A. Real-life neuroscience: an ecological approach to brain and behavior research. Perspect. Psychol. Sci. 14 (5), 841–859. https://doi.org/10.1177/1745691619856350 (2019).
Snow, J. C. & Culham, J. C. The treachery of images: how realism influences brain and behavior. Trends Cogn. Sci. 25 (6), 506–519. https://doi.org/10.1016/j.tics.2021.02.008 (2021).
Fries, P. A mechanism for cognitive dynamics: neuronal communication through neuronal coherence. Trends Cogn. Sci. 9 (10), 474–480. https://doi.org/10.1016/j.tics.2005.08.011 (2005).
Brunet, N. M. et al. Stimulus repetition modulates gamma-band synchronization in primate visual cortex. Proc. Nat Acad. Sci. 111(9), 3626–3631. (2014). https://doi.org/10.1073/pnas.1309714111
Chen, X. et al. Alpha oscillatory activity is causally linked to working memory retention. PLoS Biol. 21 (2), e3001999. https://doi.org/10.1371/journal.pbio.3001999 (2023).
Hermes, D., Miller, K. J., Wandell, B. A. & Winawer, J. Gamma oscillations in visual cortex: the stimulus matters. Trends Cogn. Sci. 19 (2), 57–58. https://doi.org/10.1016/j.tics.2014.12.009 (2015).
Hermes, D., Miller, K. J., Wandell, B. A. & Winawer, J. Stimulus dependence of gamma oscillations in human visual cortex. Cereb. Cortex. 25 (9), 2951–2959. https://doi.org/10.1093/cercor/bhu091 (2015).
Martinovic, J., Gruber, T., Hantsch, A. & Müller, M. M. Induced gamma-band activity is related to the time point of object identification. Brain Res. 1198, 93–106. https://doi.org/10.1016/j.brainres.2007.12.050 (2008).
Chen, Y. & Farivar, R. Natural scene representations in the gamma band are prototypical across subjects. NeuroImage 221, 117010. https://doi.org/10.1016/j.neuroimage.2020.117010 (2020).
Bridwell, D. A., Roth, C., Gupta, C. N. & Calhoun, V. D. Cortical response similarities predict which audiovisual clips individuals viewed, but are unrelated to clip preference. PLOS One. 10 (6), e0128833. https://doi.org/10.1371/journal.pone.0128833 (2015).
Lankinen, K., Saari, J., Hari, R. & Koskinen, M. Intersubject consistency of cortical MEG signals during movie viewing. NeuroImage 92, 217–224. https://doi.org/10.1016/j.neuroimage.2014.02.004 (2014).
Chang, W. T. et al. Combined MEG and EEG show reliable patterns of electromagnetic brain activity during natural viewing. NeuroImage 114, 49–56. https://doi.org/10.1016/j.neuroimage.2015.03.066 (2015).
Chen, L., Cichy, R. M. & Kaiser, D. Representational shifts from feedforward to feedback rhythms index phenomenological integration in naturalistic vision. Commun. Biology. 8 (1), 576. https://doi.org/10.1038/s42003-025-08011-0 (2025).
Norcia, A. M., Appelbaum, L. G., Ales, J. M., Cottereau, B. R. & Rossion, B. The steady-state visual evoked potential in vision research: a review. J. Vis. 15 (6), 4. https://doi.org/10.1167/15.6.4 (2015).
Adrian, E. D. & Matthews, B. H. C. The berger rhythm: potential changes from the occipital lobes in man. Brain 57 (4), 355–385. https://doi.org/10.1093/brain/57.4.355 (1934).
Cole, S. R. & Voytek, B. Brain oscillations and the importance of waveform shape. Trends Cogn. Sci. 21 (2), 137–149. https://doi.org/10.1016/j.tics.2016.12.008 (2017).
Dowsett, J., Dieterich, M. & Taylor, P. C. J. Mobile steady-state evoked potential recording: dissociable neural effects of real-world navigation and visual stimulation. J. Neurosci. Methods. 332, 108540. https://doi.org/10.1016/j.jneumeth.2019.108540 (2020).
Balestrieri, E. et al. Beyond oscillations—toward a richer characterization of brain states. Imaging Neurosci. 3, imag_a_00499. https://doi.org/10.1162/imag_a_00499 (2025).
Herrmann, C. S. Human EEG responses to 1-100 Hz flicker: resonance phenomena in visual cortex and their potential correlation to cognitive phenomena. Exp. Brain Res. 137 (3–4), 346–353. https://doi.org/10.1007/s002210100682 (2001).
Peelen, M. V. & Downing, P. E. Testing cognitive theories with multivariate pattern analysis of neuroimaging data. Nat. Hum. Behav. 7 (9), 1430–1441. https://doi.org/10.1038/s41562-023-01680-z (2023).
Haxby, J. V. et al. Distributed and overlapping representations of faces and objects in ventral temporal cortex. Science 293 (5539), 2425–2430. https://doi.org/10.1126/science.1063736 (2001).
Bae, G. Y. & Luck, S. J. Dissociable decoding of spatial attention and working memory from EEG oscillations and sustained potentials. J. Neuroscience: Official J. Soc. Neurosci. 38 (2), 409–422. https://doi.org/10.1523/JNEUROSCI.2860-17.2017 (2018).
Santos-Mayo, A. et al. Decoding in the fourth dimension: classification of temporal patterns and their generalization across locations. Hum. Brain. Mapp. 46 (2), e70152. https://doi.org/10.1002/hbm.70152 (2025).
Misaki, M., Kim, Y., Bandettini, P. A. & Kriegeskorte, N. Comparison of multivariate classifiers and response normalizations for pattern-information fMRI. NeuroImage 53 (1), 103–118. https://doi.org/10.1016/j.neuroimage.2010.05.051 (2010).
Jaeger, C. et al. Targeted rhythmic visual stimulation at individual participants’ intrinsic alpha frequency causes selective increase of occipitoparietal BOLD-fMRI and EEG functional connectivity. NeuroImage 270, 119981. https://doi.org/10.1016/j.neuroimage.2023.119981 (2023).
Nuttall, R. et al. Evoked responses to rhythmic visual stimulation vary across sources of intrinsic alpha activity in humans. Sci. Rep. 12 (1), 5986. https://doi.org/10.1038/s41598-022-09922-2 (2022).
Regan, D. Human brain electrophysiology: Evoked potentials and evoked magnetic fields in science and medicine (Elsevier, 1989).
McTeague, L. M., Gruss, L. F. & Keil, A. Aversive learning shapes neuronal orientation tuning in human visual cortex. Nat. Commun. 6 (1), 7823. https://doi.org/10.1038/ncomms8823 (2015).
Bashford, L. et al. The neurophysiological representation of imagined somatosensory percepts in human cortex. J. Neuroscience: Official J. Soc. Neurosci. 41 (10), 2177–2185. https://doi.org/10.1523/JNEUROSCI.2460-20.2021 (2021).
Bragin, A., Engel, J., Wilson, C. L., Fried, I. & Buzsáki, G. High-frequency oscillations in human brain. Hippocampus 9 (2), 137–142. https://doi.org/10.1002/(SICI)1098-1063(1999)9:2%3C137::AID-HIPO5%3E3.0.CO;2-0 (1999).
Cole, S. & Voytek, B. Cycle-by-cycle analysis of neural oscillations. J. Neurophysiol. 122 (2), 849–861. https://doi.org/10.1152/jn.00273.2019 (2019).
de Vries, I. E. J., Marinato, G. & Baldauf, D. Decoding object-based auditory attention from source-reconstructed MEG alpha oscillations. J. Neuroscience: Official J. Soc. Neurosci. 41 (41), 8603–8617. https://doi.org/10.1523/JNEUROSCI.0583-21.2021 (2021).
Bonnefond, M. & Jensen, O. The role of alpha oscillations in resisting distraction. Trends Cogn. Sci. 29(4), 368–379. https://doi.org/Natural (2025). scene-discriminative information was preferentially expressed in the gamma band.
Cruz, G. et al. Oscillatory brain activity in the canonical alpha-band conceals distinct mechanisms in attention. J. Neurosci. 45 (1), e0918242024. https://doi.org/10.1523/JNEUROSCI.0918-24.2024 (2025).
Benwell, C. S. Y., Coldea, A., Harvey, M. & Thut, G. Low pre-stimulus EEG alpha power amplifies visual awareness but not visual sensitivity. Eur. J. Neurosci. 55 (11–12), 3125–3140. https://doi.org/10.1111/ejn.15166 (2022).
Peylo, C., Hilla, Y. & Sauseng, P. Cause or consequence? Alpha oscillations in visuospatial attention. Trends Neurosci. 44 (9), 705–713. https://doi.org/10.1016/j.tins.2021.05.004 (2021).
Baldauf, D. & Desimone, R. Neural mechanisms of object-based attention. Science 344 (6182), 424–427. https://doi.org/10.1126/science.1247003 (2014).
Mazzoni, A., Brunel, N., Cavallari, S., Logothetis, N. K. & Panzeri, S. Cortical dynamics during naturalistic sensory stimulations: experiments and models. J. Physiology-Paris. 105 (1–3), 2–15. https://doi.org/10.1016/j.jphysparis.2011.07.014 (2011).
van Kerkoerle, T. et al. Alpha and gamma oscillations characterize feedback and feedforward processing in monkey visual cortex. Proc. Natl. Acad. Sci. 111 (40), 14332–14341. https://doi.org/10.1073/pnas.1402773111 (2014).
Keitel, A. et al. Brain rhythms in cognition—Controversies and future directions (ArXiv:2507.15639). arXiv. https://doi.org/10.48550/arXiv.2507.15639 (2025).
Spaak, E., Bonnefond, M., Maier, A., Leopold, D. A. & Jensen, O. Layer-specific entrainment of γ-band neural activity by the α rhythm in monkey visual cortex. Curr. Biology: CB. 22 (24), 2313–2318. https://doi.org/10.1016/j.cub.2012.10.020 (2012).
Ronconi, L., Balestrieri, E., Baldauf, D. & Melcher, D. Distinct cortical networks subserve spatio-temporal sampling in vision through different oscillatory rhythms. J. Cogn. Neurosci. 36 (4), 572–589. https://doi.org/10.1162/jocn_a_02006 (2024).
Mathewson, K. E., Gratton, G., Fabiani, M., Beck, D. M. & Ro, T. To see or not to see: prestimulus alpha phase predicts visual awareness. J. Neuroscience: Official J. Soc. Neurosci. 29 (9), 2725–2732. https://doi.org/10.1523/JNEUROSCI.3963-08.2009 (2009).
Jensen, O., Bonnefond, M. & VanRullen, R. An oscillatory mechanism for prioritizing salient unattended stimuli. Trends Cogn. Sci. 16 (4), 200–206. https://doi.org/10.1016/j.tics.2012.03.002 (2012).
Bagherzadeh, Y., Baldauf, D., Pantazis, D. & Desimone, R. Alpha synchrony and the neurofeedback control of spatial attention. Neuron 105 (3), 577–587e5. https://doi.org/10.1016/j.neuron.2019.11.001 (2020).
de Vries, E. & van Ede, F. Microsaccades reveal preserved spatial organisation in visual working memory despite decay in location-based rehearsal. Cognition 259, 106111. https://doi.org/10.1016/j.cognition.2025.106111 (2025).
Bae, G. Y. & Luck, S. J. Reactivation of previous experiences in a working memory task. Psychol. Sci. 30 (4), 587–595. https://doi.org/10.1177/0956797619830398 (2019).
Saha, S. et al. Progress in brain computer interface: challenges and opportunities. Front. Syst. Neurosci. 15, 578875. https://doi.org/10.3389/fnsys.2021.578875 (2021).
Wang, G. et al. Human-centred physical neuromorphics with visual brain-computer interfaces. Nat. Commun. 15 (1), 6393. https://doi.org/10.1038/s41467-024-50775-2 (2024).
Ferrante, M., Boccato, T., Ozcelik, F., VanRullen, R. & Toschi, N. Through their eyes: multi-subject brain decoding with simple alignment techniques. Imaging Neurosci. 2, imag–2. https://doi.org/10.1162/imag_a_00170 (2024).
Trammel, T., Khodayari, N., Luck, S. J., Traxler, M. J. & Swaab, T. Y. Decoding semantic relatedness and prediction from EEG: A classification method comparison. NeuroImage 277, 120268. https://doi.org/10.1016/j.neuroimage.2023.120268 (2023).
Cirulli, F., Spencer, S. J. & Zhang, C. Diversity matters. Neuroscience 564, 319–321. https://doi.org/10.1016/j.neuroscience.2024.11.057 (2025).
Shen, F. X. et al. Emerging ethical issues raised by highly portable MRI research in remote and resource-limited international settings. NeuroImage 238, 118210. https://doi.org/10.1016/j.neuroimage.2021.118210 (2021).
Debener, S., Emkes, R., De Vos, M. & Bleichner, M. Unobtrusive ambulatory EEG using a smartphone and flexible printed electrodes around the ear. Sci. Rep. 5 (1), 16743. https://doi.org/10.1038/srep16743 (2015).
Notbohm, A., Kurths, J. & Herrmann, C. S. Modification of brain oscillations via rhythmic light stimulation provides evidence for entrainment but not for superposition of event-related responses. Front. Hum. Neurosci. https://doi.org/10.3389/fnhum.2016.00010 (2016).
Parkes, L. M., Fries, P., Kerskens, C. M. & Norris, D. G. Reduced BOLD response to periodic visual stimulation. NeuroImage 21 (1), 236–243. https://doi.org/10.1016/j.neuroimage.2003.08.025 (2004).
Buracas, G. T., Zador, A. M., DeWeese, M. R. & Albright, T. D. Efficient discrimination of temporal patterns by motion-sensitive neurons in primate visual cortex. Neuron 20 (5), 959–969. https://doi.org/10.1016/s0896-6273(00)80477-8 (1998).
Tabarelli, D., Keitel, C., Gross, J. & Baldauf, D. Spatial attention enhances cortical tracking of quasi-rhythmic visual stimuli. NeuroImage 208, 116444. https://doi.org/10.1016/j.neuroimage.2019.116444 (2020).
Dowsett, J., Herrmann, C. S., Dieterich, M. & Taylor, P. C. J. Shift in lateralization during illusory self-motion: EEG responses to visual flicker at 10 Hz and frequency‐specific modulation by tACS. Eur. J. Neurosci. 51 (7), 1657–1675. https://doi.org/10.1111/ejn.14543 (2020).
Cohen, M. X. (2017). MATLAB for Brain and CognitiveScientists. MIT Press.
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JD contributed to all aspects of the manuscript.IMM contributed to data collection, writing analysis code and writing of the manuscript.PT contributed to the design of the experiment and writing/editing of the manuscript.
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Dowsett, J., Muñoz, I.M. & Taylor, P. Decoding real-world visual scenes from alpha and gamma band flicker evoked oscillations in human EEG. Sci Rep (2026). https://doi.org/10.1038/s41598-026-42197-5
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DOI: https://doi.org/10.1038/s41598-026-42197-5