Abstract
Dynamic ocean features (e.g., currents, eddies) can drive large-scale transport of water masses and nutrients, facilitating localized plankton blooms and affecting the migration patterns of higher trophic levels. The whale shark, Rhincodon typus, is a highly migratory planktivore known to aggregate in areas of ephemeral food abundance. While the species’ presence has been widely correlated with environmental variables such as water temperature and chlorophyll-a, the Red Sea population appears to be primarily associated with other oceanographic variables. Here, we investigated factors influencing juvenile whale shark presence within the Red Sea by correlating previously published tracking data with remote sensing measurements of environmental variables using Generalized Additive Mixed Models (GAMM). Model results revealed a significant correlation between whale shark presence and mixed-layer depth, wind direction, north-south current velocity, and temperature. The model trends and satellite-tracking analysis also indicated that whale sharks would spend more time actively following eddies within the basin. This new insight can help understand how whale sharks rely on ocean dynamics in this nutrient-poor subtropical basin, as regions influenced by wind, chlorophyll enrichment, and vertical mixing offer increased foraging opportunities for the species. It may also serve as a reference point for future research that identifies key whale shark habitats and considers the impact of climate change on preferred environments.
Data availability
All data generated or analyzed in this study are available from the corresponding author upon reasonable request.
Abbreviations
- IUCN:
-
International Union for Conservation of Nature
- CITES:
-
Convention on International Trade in Endangered Species
- SPOT:
-
Satellite Positioning and Temperature Tag
- GAMM:
-
Generalized Additive Mixed Model
- BRAN:
-
Bluelink Reanalysis Model
- SST:
-
Sea Surface Temperature
- SSH:
-
Sea Surface Height
- MLD:
-
Mixed Layer Depth
- Wspd:
-
Wind Speed
- Wdir:
-
Wind Direction
- VCS:
-
Vertical current speed
- HCS:
-
Horizontal current speed
- VCUR:
-
North South Current Velocity
- UCUR:
-
West East current
- WCUR:
-
Bottom surface current
- ChlA:
-
Chlorophyll-a
- AIC:
-
Akaike Information Criterion
- ANOVA:
-
Analysis of Variance
- GAIW:
-
Gulf of Aden Intermediate Waters
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Acknowledgements
The authors are grateful to Djidenou Montcho for the assistance and opinions on the model development, João Curdia for idea conceptualization and preliminary statistical trials, and Ronald Cadiz for help with figure creation. We thank Kelly Quinn for her approval of the use of her whale shark illustration on our figures. We are also grateful to all the people who supported us throughout the development of this study. Finally, we would like to thank Ana Bigio (Scientific Illustrator, Research Communication - KAUST) for her amazing schematic work on Figure 7.
Funding
This work was financially supported by KAUST baseline research funds (to MLB and BHJ), and previous whale shark tracking data were also provided by KAUST award nos. USA00002 and KSA 00011 (to SRT), and the United States National Science Foundation (OCE 0825148 to SRT and GBS). The funders had no role in study design, data collection and analysis, decision to publish, or manuscript preparation.
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Raquel L. Ostrovski: Conceptualization, Methodology, Software, Validation, Formal analysis, Investigation, Data curation, Writing - original draft, Writing - review and editing, Visualization; Jesse E. M. Cochran: Conceptualization, Data curation, Investigation, Supervision, Visualization, Writing - review and editing; Yuri Niella: Data curation, Formal analysis, Investigation, Methodology, Software, Writing - original draft; Alkiviadis Kalampokis: Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Supervision, Visualization, Writing - review and editing; Israel J. S. Filho: Data curation, Formal analysis, Methodology, Software; Ute Langner: Investigation, Software, Visualization; Royale S. Hardenstine: Investigation, Writing- review and editing; Paula Moraga: Methodology, Validation, Visualization; Michael Berumen: Data curation, Funding acquisition, Resources, Validation, Visualization, Writing - review and editing; Burton Jones: Conceptualization, Funding acquisition, Investigation, Project administration, Resources, Supervision, Validation, Visualization, Writing - review and editing.
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Ostrovski, R.L., Cochran, J.E., Niella, Y. et al. Flow-driven patterns of whale shark movement in the Red Sea. Sci Rep (2026). https://doi.org/10.1038/s41598-026-45029-8
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DOI: https://doi.org/10.1038/s41598-026-45029-8