Abstract
Ricklefs replies - Purvis1 states that, under random gradual change, clades accumulate variance in proportion to their total branch length. Accordingly, at a given age, clades with more species should exhibit greater variance. This is incorrect, as shown both analytically2,3 and by simulation4,5, and the error underlines a misunderstanding of my analysis.
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Ricklefs, R. How do characters evolve? (reply). Nature 432, 166 (2004). https://doi.org/10.1038/nature03093
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DOI: https://doi.org/10.1038/nature03093
