Thank you for visiting nature.com. You are using a browser version with limited support for CSS. To obtain
the best experience, we recommend you use a more up to date browser (or turn off compatibility mode in
Internet Explorer). In the meantime, to ensure continued support, we are displaying the site without styles
and JavaScript.
And the locations of study sites mentioned in this text include: Sanbao cave8, Xiangride (XRD)55, Gonghai (GH)56, and Dongge cave4. The base map was generated using ArcGIS software.
