Fig. 3: Relationships of spatial dissimilarity of endemic alpine species with paleo fluctuation in upper range limit of trees, paleotemperature, and paleo precipitation since the Last Glacial Period (LGM) across all regions over the Tibetan Plateau.

The relationships of (a–c) beta-diversity (β-diversity), (d–f) species replacement and (g–i) nestedness to paleo fluctuation in upper range limit of trees (gray circles), growing-season temperature (red circles), and precipitation (blue circles) since LGM across all regions over the Tibetan Plateau. Here the region is defined to be consisted of a target county and its surrounding counties. The σ represents the paleo fluctuation for each variable and is calculated as the standard deviation of its high-frequency component based on the Empirical Mode Decomposition method. The black lines were fitted with linear regressions. The asterisks indicate statistical significance by two-tailed Student’s t-test (P < 0.01).