Table 1 Morphological characteristics of knotweed species based on literature review3,11,12,13.
Feature | R. japonica Houtt. (Japanese Knotweed) | R. x bohemica Chrtek & Chrtková (Bohemian Knotweed) | R. sachalinensis (F. Schmidt) Nakai (Giant Knotweed) |
|---|---|---|---|
Leaf shape and size | Broadly ovate-triangular, 5–18 cm long, 4–13 cm wide; typically up to 15 cm long and 10 cm wide | Intermediate between R. japonica and R. sachalinensis; typically broadly ovate, 10–27 cm long, 9–22 cm wide; typically up to 23 cm long and 20 cm wide | Broadly oblong-ovate, heart-shaped or elongated, 15–43 cm long and 10-27 cm wide; typically up to 40 cm long and 25 cm wide length: width ratio c. 1.5 |
Leaf base | Acuminate/ ovate/ nearly truncate/ truncate/ bluntly wedge-shaped narrowed | Sagittate or moderately cordate; typically base bluntly wedge-shaped (upper leaves) or slightly cordate (lower leaves) | Deeply cordate |
Leaf apex/tip | Pointed, cuspidate or acuminate, narrowed into a long tip, curved intermediate | Intermediate, variable pointed, slightly acuminate | Acute, tapering to obtuse/ rounded/ bluntly pointed |
Leaf trichomes | Sparse (few to none), short, on leaf underside along veins, reduced to short, single-celled papillae with a strongly swollen base mostly glabrous or scabrous to tuburculate (type A) | Sparse, intermediate trichome shape, density, and length; short, 1-4-celled, with a strongly thickened base (type B) | Dense, longer hairs on leaf underside, 1–5 cells, and 8 or greater (type C) |
Stem features | Hollow, green with reddish/purplish/brown speckles and reddish nodes, bamboo-like, up to 2–3 m tall; grow in groups of several to a dozen from a rhizome node (shoot clump) | Variable, often faintly speckled, intermediate height, typically up to 3,5 m; grow in groups of several to a dozen from a rhizome node (shoot clump) | Hollow, robust, usually without speckles, up to 4–4,5 m tall; grow singly or in groups from a rhizome node (shoot clump) |
Growth habit | Dense, bamboo-like thickets, upright growth | Combines the dense formation of R. japonica with the spreading nature of R. sachalinensis | Less dense thickets, spreading growth habit |